Clerodendrum quadriloculare (Blanco) Merr., Verbenaceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Risk assessment results:  High risk, score: 11 (Go to the risk assessment)

Other Latin names:  Ligustrum quadriloculare Blanco

Common name(s): [more details]

English: bronze-leaved clerodendrum, fire works, Philippine glorybower

Marshallese: tirooj in belle

Other: bagauac

Palauan: kleuang

Pohnpeian: tuhkehn Palau

Samoan: losa, losa, losa Fiti, losa Fiti

Habit:  shrub

Description:  "An erect, glabrous or nearly glabrous shrub or small tree 2 to 5 m high.  Leaves oblong, 15-20 cm long, apex shortly acuminate, base rounded, the upper surface green, the lower surface usually uniformly dark-purple.  Cymes terminal, panicled, usually many-flowered.  Calyx urceolate, purple about 1 cm long, about 2 mm in diameter, the limb spreading, the lobes oblong-elliptic, about 1.5 cm long.  Fruit ellipsoid, 1 to 1.5 cm long, purplish, the persistent calyx red and 1 to 1.5 cm long"  (Merrill, 1912; p. 402).

"Shrub from the Philippines with large, paired oval leaves that are dark green above and reddish purple below. The flowers, which are produced in very showy large clusters, each have a narrow pink tube that is commonly 3 inches or more in length, ending in five white, spreading lobes" (University of Hawai'i Botany Department).

Habitat/ecology:  Commonly planted as an ornamental for its foliage and showy flowers.

Propagation:  Produces large amounts of seed and suckers profusely from the roots.

Native range:  New Guinea, Philippines (GRIN).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Manu‘a Islands	Ta‘u Island	introduced cultivated	Space, James C./Flynn, Tim (2000) (pp. 6, 13)
American Samoa Tutuila Islands	Tutuila Island	introduced cultivated	Space, James C./Flynn, Tim (2000) (p. 6)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Tinian Island	introduced cultivated	Space, James C./Falanruw, Marjorie (1999) (p. 10)
Federated States of Micronesia Chuuk Islands	Fefan Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (p. 5) Voucher: BMW 5804 (BISH, BRI, GUAM)
Federated States of Micronesia Chuuk Islands	Fefan Island		Bishop Museum (Honolulu) (2000) (voucher ID: BISH 687758) Taxon name on voucher: Clerodendrum quadriloculare (Blanco) Merr.
Federated States of Micronesia Chuuk Islands	Fefan Island		Bishop Museum (Honolulu) (2000) (voucher ID: BISH 687759) Taxon name on voucher: Clerodendrum quadriloculare (Blanco) Merr.
Federated States of Micronesia Chuuk Islands	Tol Island	introduced cultivated	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (p. 5)
Federated States of Micronesia Chuuk Islands	Weno (Moen) Island	introduced cultivated	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (p. 5)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced cultivated	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Waguk, Erick E. (2000) (p. 5)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive cultivated	Lorence, David H./Flynn, Tim (2005) (p. 16)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive	Josekutty, P. C./Wakuk, E. E./Joseph, M. J. (2002) (p. 62)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Swarbrick, John T. (1997) (p. 90)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive	Meyer, Jean-Yves (2000) (p. 90)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive cultivated	Space, James C./Falanruw, Marjorie (1999) (pp. 4, 10)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive cultivated	Lorence, David H./Flynn, Tim (1998) (p. 31) Becoming naturalized
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	introduced cultivated	Space, James C./Falanruw, Marjorie (1999) (p. 10)
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235)
Guam Guam Island	Guam Island	introduced	Meyer, Jean-Yves (2000) (p. 95) "Potential invader".
Guam Guam Island	Guam Island	introduced cultivated	Space, James C./Falanruw, Marjorie (1999) (p. 10)
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1966) (voucher ID: BISH 652423) Taxon name on voucher: Clerodendrum quadriloculare (Blanco) Merr.
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1963) (voucher ID: BISH 652444) Taxon name on voucher: Clerodendrum quadriloculare (Blanco) Merr.
State of Hawaii Hawaiian Islands	Hawaiian Islands	introduced cultivated	Staples, George W./Herbst, Derral/Imada, Clyde T. (2000) (p. 31)
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll	introduced cultivated	Whistler, W. A./Steele, O. (1999) (p. 103)
Marshall Islands Ratak Chain	Majuro (Mãjro) Atoll	introduced cultivated	Vander Velde, Nancy (2003) (p. 131)
Palau Palau (main island group)	Angaur Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 88)
Palau Palau (main island group)	Angaur Island	introduced invasive cultivated	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 10)
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 84)
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 10)
Palau Palau (main island group)	Kayangel (Ngcheangel) Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 89)
Palau Palau (main island group)	Koror Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 80)
Palau Palau (main island group)	Koror Island	introduced cultivated	Lorence, David H./Flynn, Tim (1998) (p. 39)
Palau Palau (main island group)	Koror Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 611196) Taxon name on voucher: Clerodendrum quadriloculare (Blanco) Merr.
Palau Palau (main island group)	Koror Island		Beleu National Museum (Republic of Palau. Koror.) (1991) (voucher ID: BNM 9574) Taxon name on voucher: Clerodendrum quadriloculare (Blanco) Merrill
Palau Palau (main island group)	Ngercheu Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 92) Carp Resort
Palau Palau (main island group)	Ngerkebesang Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 80)
Palau Palau (main island group)	Ngerkebesang Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 10)
Palau Palau (main island group)	Peleliu Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 7-8, 90)
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)		U.S. Dept. Agr., Agr. Res. Serv. (2009)
Philippines Philippine Islands	Philippine Islands	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Samoa Western Samoa Islands	Savai‘i Island	introduced invasive cultivated	Space, James C./Flynn, Tim (2002) (p. 6)
Samoa Western Samoa Islands	Upolu Island	introduced invasive cultivated	Space, James C./Flynn, Tim (2002) (p. 6)

Comments:  Suckers and sprouts profusely, making it an undesirable tree to plant for ornamental and other purposes. Mostly found along roads and cultivated in yards. However, On Pohnpei it was seen growing in full shade under the forest canopy in a dense, monospecific understory (Space & Falanruw, 1999). Planting of this species should be discouraged and it should be monitored closely for spread.

Additional information:
Additional online information about Clerodendrum quadriloculare is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Clerodendrum quadriloculare as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Clerodendrum quadriloculare may be available from the Germplasm Resources Information Network (GRIN).

Other Latin names:  Ligustrum quadriloculare Blanco

References:

Beleu National Museum (Republic of Palau. Koror.). 1991. Voucher specimen #BNM9574(Herbst, Stemmermann & Canfield 9574).

Bishop Museum (Honolulu). 1963. Voucher specimen #BISH 652444 (Fosberg, F.R. 43480).

Bishop Museum (Honolulu). 1966. Voucher specimen #BISH 652423 (Evans, M. 1639).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 611196 (Herbst, D.R. 9574).

Bishop Museum (Honolulu). 2000. Voucher specimen #BISH 687758 (Waterhouse, B.M. BMW5804).

Bishop Museum (Honolulu). 2000. Voucher specimen #BISH 687759 (Waterhouse, B.M. BMW5804).

Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.

Josekutty, P. C./Wakuk, E. E./Joseph, M. J. 2002. Invasive/weedy angiosperms in Kosrae, Federated States of Micronesia. In: Invasive species and their management. Micronesica Suppl. 6:61-65.

Lorence, David H./Flynn, Tim. 1998. Checklist of the plants of Palau. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawai‘i. 40 pp.

Lorence, David H./Flynn, Tim. 1998. Checklist of the plants of Pohnpei. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawai‘i. 21 pp.

Lorence, David H./Flynn, Tim. 2005. Checklist of the plants of Kosrae. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawai‘i. 23 pp.

Merrill, E. D. 1912. A flora of Manila. Manila. 490 pp. (1968 reprint).

Meyer, Jean-Yves. 2000. Preliminary review of the invasive plants in the Pacific islands (SPREP Member Countries). In: Sherley, G. (tech. ed.). Invasive species in the Pacific: A technical review and draft regional strategy. South Pacific Regional Environment Programme, Samoa. 190 pp.

Space, James C./Falanruw, Marjorie. 1999. Observations on invasive plant species in Micronesia. USDA Forest Service, Honolulu. Report to the Pacific Islands Committee, Council of Western State Foresters. USDA Forest Service, Honolulu. 32 pp.

Space, James C./Flynn, Tim. 2000. Observations on invasive plant species in American Samoa.  USDA Forest Service, Honolulu. 51 pp.

Space, James C./Flynn, Tim. 2002. Report to the Government of Samoa on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 83 pp.

Space, James C./Lorence, David H./LaRosa, Anne Marie. 2009. Report to the Republic of Palau: 2008 update on Invasive Plant Species. USDA Forest Service, Hilo, Hawai‘i. (In press).

Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. 2000. Invasive plant species in Chuuk, Federated States of Micronesia.  USDA Forest Service, Honolulu. 41 pp.

Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Waguk, Erick E. 2000. Invasive plant species on Kosrae, Federated States of Micronesia.  USDA Forest Service, Honolulu. 43 pp.

Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar. 2003. Report to the Republic of Palau on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 174 pp.

Staples, George W./Herbst, Derral/Imada, Clyde T. 2000. Survey of invasive or potentially invasive cultivated plants in Hawai‘i. Bishop Museum Occasional Papers No. 65. 35 pp.

Swarbrick, John T. 1997. Weeds of the Pacific Islands. Technical paper no. 209. South Pacific Commission, Noumea, New Caledonia. 124 pp.

U.S. Dept. Agr., Agr. Res. Serv. 2009. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.

Vander Velde, Nancy. 2003. The vascular plants of Majuro Atoll, Republic of the Marshall Islands. Smithsonian Institution, Atoll Research Bulletin No. 503:1-141.

Whistler, W. A. 2000. Tropical ornamentals: a guide. Timber Press, Portland, Oregon. 542 pp.

Whistler, W. A./Steele, O. 1999. Botanical survey of the United States of America Kwajalein Atoll (USAKA) Islands. Prepared for Oak Ridge Institute for Science and Education and the U. S. Army Environmental Center. 111 pp.