Urochloa mutica (Forssk.) T.Q. Nguyen, Poaceae

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Present on Pacific Islands?  yes

Threat only at high elevations?  no

Risk assessment results: 

Reject, score: 14 (Go to the risk assessment (Australia))
High risk, score: 12 (Go to the risk assessment (Pacific))

Other Latin names:  Brachiaria mutica (Forssk.) Stapf, Panicum muticum Forssk., Panicum purpurascens Raddi, Panicum barbinode Trin., Panicum guadeloupense Steudel, Brachiaria purpurascens Henr.

Common name(s): [more details]

English: buffalo grass, California grass, Dutch grass, giant couch, Mauritius grass, Para grass, Scotch grass, tall panicum, water grass

French: herbe borer, herbe de Guinée, herbe de Para

Maori (Cook Islands): mauku puakatoro

Spanish: gramalote, hierba de Pará, pasto Pará, zacuti Para

Tongan: puakatau

Habit:  grass

Description:  "Perennial; culms decumbent and rooting at base, 2-5 m long, the nodes densely villous; sheaths mostly longer than internodes, papillose or papillose-pilose, densely short-puberulent on collar; ligule a row of hairs about 1 mm long; blades 10-30 cm long, 10-15 mm wide, acuminate, glabrous on both surfaces, margins scabrous; inflorescences 10-25 cm long, about half as broad, lowermost branches compound, upper simple, branchlets and upper branches bearing spikelets solitary or in 2's or 3's on 1 side of a flattened rachis, this, like main axis, angled, scabrous on angles; spikelets glabrous, pointed, about 3 mm long; first glume acute, about 1 mm long, not clasping at base; second glume and sterile lemma equal, the latter subtending a hyaline palea; fertile lemma slightly shorter, obtuse, minutely transversely rugose"  (Wiggins & Porter, 1971; pp. 868-869).

"Culms decumbent, rooting at the base, up to 2.4 m high, with densely hairy nodes, hairy sheaths, blades 10-30 cm long and 13 mm wide, glabrous.  Panicle up to 30 cm long, with numerous (about 10) spreading branches, 2.5-10 cm long; spikelets up to 4.5 mm long, elliptic, sometimes purplish rachis more or less flat, herbaceous, ribbon-like.  First glume 1/3-1/2 as long as the spikelet.  Fruit with minute transverse grooves (Stone, 1970; pp. 203-204).

Description from World Grass Species.

Habitat/ecology:  "Wet fields, ditches and gullies between sea level and 700 m. This perennial grass can reach heights of 2 m. It forms dense monotypic stands by layering from trailing stems. It will overgrow most shrubs and trees in its habitat. It has mild allelopathic activity (Chou and Young 1975).  Fire is rare in its habitat but the dense stands rapidly regenerate from any damage that they suffer" (Smith, 1985; p. 186). 

Invasive to 5000 feet in New Guinea (Henty, 1969; p. 39).  In Hawai‘i, "naturalized and usually forming dense mats in areas of high soil moisture or nearby open water, such as along stream banks, drainage ditches, and roadsides, as well as other disturbed sites, 0-1,060 m" (Wagner et al., 1999; p. 1504).  In Fiji, "widespread from sea level to about 800 m, especially in the wet zones, where it occurs in pastures and cultivated fields, along roadsides, on river banks, in swamps, in clearings, and along forest trails" (Smith, 1979; p. 329). In New Caledonia, "maintenant largement dispersé et souvent abondant en stations plus ou moins humides" (MacKee, 1994; p. 57).

"Interferes with stream flow and poses a nuisance to marine navigation when rafts of the grass float out to sea. Forms monotypic stands in forest openings and marshes, displacing native plants and destroying bird habitats."  (Motooka et al., 2003). "In moist or wet ground, sometimes growing in water, throughout tropical and subtropical America, usually at low altitudes; cultivated for forage" (Wiggins & Porter, 1971; pp. 868-869).  In Australia, "wet or seasonally flooded areas in disturbed sites including flood plains, drainage lines and creek banks.  Can form dense floating mats 1-2 m thick over water bodies.  Can tolerate water depths up to 1 m.  A highly invasive grass that spreads vigorously, choking out native species and hindering re-establishment of more desirable species.  Will tolerate brackish water and is relatively shade tolerant." (Smith, 2002; p. 80).

Propagation:  Man is the principal dispersal agent (as a pasture grass). Spreads by seed, runners and cuttings; rarely flowers. "Also by floodwaters and birds" (Smith, 2002; p. 80).

Native range:  "Probably a native of Africa, but first described from Brazilian specimens.  It is now cultivated rather widely"  (Stone, 1970; pp. 203-204).  "A native or northern Africa, this grass was probably established in tropical America in the early days of trading" (Smith, 1979; p. 329).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Manu‘a Islands	Ta‘u Island	introduced invasive	Space, James C./Flynn, Tim (2000) (p. 44)
American Samoa Tutuila Islands	Tutuila Island	introduced invasive	Space, James C./Flynn, Tim (2000) (p. 32)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 52)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island		Bishop Museum (Honolulu) (1986) (voucher ID: BISH 572382) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island		Bishop Museum (Honolulu) (1950) (voucher ID: BISH 588367) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island		Bishop Museum (Honolulu) (1946) (voucher ID: BISH 588986) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Cook Islands Southern Cook Islands	Aitutaki Atoll		McCormack, Gerald (2007)
Cook Islands Southern Cook Islands	Miti‘aro Island		McCormack, Gerald (2007)
Cook Islands Southern Cook Islands	Rarotonga Island		McCormack, Gerald (2007)
Cook Islands Southern Cook Islands	Rarotonga Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 664868) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Ecuador (Galápagos Islands) Floreana Group	Floreana Island	introduced invasive cultivated	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Isabela Group	Volcan Sierra Negra	introduced invasive cultivated	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) San Cristóbal Group	San Cristóbal Island	introduced invasive cultivated	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Santa Cruz Group	Santa Cruz Island	introduced invasive cultivated	Charles Darwin Research Station (2005)
Fiji Fiji Islands	Mbatiki Island	introduced invasive cultivated	Smith, Albert C. (1979) (p. 329)
Fiji Fiji Islands	Vanua Levu Island	introduced invasive cultivated	Smith, Albert C. (1979) (p. 329) Voucher cited: DA 8751
Fiji Fiji Islands	Vanua Levu Island		Bishop Museum (Honolulu) (1954) (voucher ID: BISH 31655) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island	introduced invasive cultivated	Smith, Albert C. (1979) (p. 329) Vouchers cited: DA 8212, DA 8251, DA 8572, DA 8663, DA 8094, Meebold 17097, DA 8428, Gillespie 3461, Bryan 216, DA 3502, DA 3517, DA 8620
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1955) (voucher ID: BISH 31656) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1924) (voucher ID: BISH 700002) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1927) (voucher ID: BISH 700003) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1932) (voucher ID: BISH 700004) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1927) (voucher ID: BISH 700005) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1924) (voucher ID: BISH 700006) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1945) (voucher ID: BISH 700007) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1947) (voucher ID: BISH 700009) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island		Bishop Museum (Honolulu) (year unknown) (voucher ID: BISH 540436) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Welsh, S. L. (1998) (p. 330) Vouchers cited: Grant 4170, Florence 3508, Fosberg & Sachet 63226
French Polynesia Society Islands	Raiatea (Havai) Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491711) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
French Polynesia Society Islands	Raiatea (Havai) Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 588738) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
French Polynesia Society Islands	Tahiti Island	introduced invasive	Welsh, S. L. (1998) (p. 330) Vouchers cited: Grant 4170, Florence 2835, Florence 2885
French Polynesia Society Islands	Tahiti Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491185) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
French Polynesia Society Islands	Tahiti Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491726) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
French Polynesia Society Islands	Tahiti Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 492086) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island	introduced cultivated	Stone, Benjamin C. (1970) (pp. 203-204) Voucher cited: Stone 4095 (GUAM)
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 52)
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1918) (voucher ID: BISH 125937) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1911) (voucher ID: BISH 125938) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1918) (voucher ID: BISH 125940) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1964) (voucher ID: BISH 125941) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1966) (voucher ID: BISH 588393) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1954) (voucher ID: BISH 588985) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1983) (voucher ID: BISH 649487) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1953) (voucher ID: BISH 699999) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1983) (voucher ID: BISH 1000209) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
State of Hawaii Hawaiian Islands	Hawai‘i (Big) Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1504)
State of Hawaii Hawaiian Islands	Kaua‘i Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1504)
State of Hawaii Hawaiian Islands	Lāna‘i Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1504)
State of Hawaii Hawaiian Islands	Maui Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1504)
State of Hawaii Hawaiian Islands	Moloka‘i Island	introduced invasive	Oppenheimer, Hank L. (2003) (p. 20) Voucher cited: Oppenheimer H110123 (BISH, PTBG)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1504)
Kiribati Line Islands	Kiritimati (Christmas) Atoll		? (year unknown)
Marshall Islands Ralik Chain	Enewetak (Ãnewetak) Atoll		Bishop Museum (Honolulu) (1976) (voucher ID: BISH 410766) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Marshall Islands Ratak Chain	Arno Atoll	introduced invasive	Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 3) "Uncommon; ruderal areas".
Marshall Islands Ratak Chain	Maloelap Atoll		Vander Velde, Nancy (year unknown)
New Caledonia New Caledonia	New Caledonia Islands	introduced invasive	Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 380) Largement dispersée et souvent abondante en stations plus ou moins humides.
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	MacKee, H. S. (1994) (p. 57) Voucher cited: cribs 832
New Zealand (offshore islands) Kermadec Islands	Raoul Island	introduced invasive	Sykes, W. R. (1977) (p. 163)
New Zealand (offshore islands) Kermadec Islands	Raoul Island	introduced invasive	Sykes, W. R./West, C. J. (1996) (p. 461)
New Zealand (offshore islands) Kermadec Islands	Raoul Island	introduced invasive	Edgar, E./Connor, H. (2000) (p. 594) "Long-abandoned orchard on terraces above The Farm".
Niue Niue	Niue Island	introduced	Sykes, W. R. (1970) (p. 234) Vouchers cited: CHR 168778, CHR 168807, CHR 168808
Niue Niue	Niue Island	introduced invasive	Whistler, W. A. (1988) (p. 46)
Niue Niue	Niue Island	introduced cultivated	Yuncker, T. G. (1943) (p. 23) Voucher cited: Yuncker 9659 (BISH) "Cultivated as a forage grass".
Niue Niue	Niue Island	introduced invasive	Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 48)
Palau Palau (main island group)	Babeldaob Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 52)
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 87)
Palau Palau (main island group)	Koror Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 52)
Palau Palau (main island group)	Koror Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 83)
Palau Palau (main island group)	Malakal Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 83)
Palau Palau (main island group)	Palau Islands (main island group)		Bishop Museum (Honolulu) (1978) (voucher ID: BISH 432334) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Palau Palau (main island group)	Palau Islands (main island group)		Bishop Museum (Honolulu) (1978) (voucher ID: BISH 432407) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)		Henty, E. E. (1969)
Philippines Philippine Islands	Philippine Islands	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)
Samoa Western Samoa Islands	Upolu Island		Bishop Museum (Honolulu) (1973) (voucher ID: BISH 417774) Taxon name on voucher: Brachiaria mutica (Forssk.) Stapf
Samoa Western Samoa Islands	Western Samoa Islands		Swarbrick, John T. (1997) (p. 89)
Solomon Islands Solomon Islands	Solomon Islands		Swarbrick, John T. (1997) (p. 89)
Solomon Islands Solomon Islands	Solomon Islands	native	Hancock, I. R./Henderson, C. P. (1988) (p. 52)
Tonga Ha‘apai Group	Lifuka and Foa Islands	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 71)
Tonga Tongatapu Group	‘Eua Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 58)
Tonga Tongatapu Group	Tongatapu Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 51)
Tonga Tongatapu Group	Tongatapu Island		Whistler, W. A. (1992) (p. 97) Vouchers cited: Soakai 644, Soakai 786
Tonga Vava‘u Group	Vava‘u Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 65)
United States (other Pacific offshore islands) Midway Atoll	Midway Atoll	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1504)
Vanuatu New Hebrides Islands	Vanuatu (Republic of)		Swarbrick, John T. (1997) (p. 89)
Wallis and Futuna Wallis and Futuna (Horne) Islands	Wallis and Futuna Islands		Whistler, W. A. (1988) (p. 46)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia Australia (continental)	Northern Territory	introduced invasive	Smith, Nicholas M. (2002) (p. 80)
Australia Australia (continental)	Queensland	introduced invasive	Smith, Nicholas M. (2002) (p. 80)
Cambodia Cambodia	Cambodia (Kingdom of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)
Colombia Colombia	Colombia (Republic of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 193-195)
Costa Rica Costa Rica	Costa Rica (Republic of)		Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 52)
Ecuador (Mainland) Ecuador	Ecuador (Republic of) (continental)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 193-195)
Indonesia Indonesia	Indonesia (Republic of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)
Malaysia Malaysia	Malaysia (country of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)
Mexico Mexico	Mexico (United Mexican States)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 193-195)
Perú Perú	Perú (Republic of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 193-195)
Singapore Singapore	Singapore (Republic of)	introduced invasive	Waterhouse, D. F. (1993) (pp. 63, 76)
Taiwan Taiwan	Taiwan Island	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)
Thailand Thailand	Thailand (Kingdom of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)
Vietnam Vietnam	Vietnam (Socialist Republic of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 191-194)

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia (Indian Ocean offshore islands) Christmas Island Group	Christmas Island	introduced invasive	Swarbrick, J. T. (1997) (p. 106) Naturalized
La Réunion (France) La Réunion Island	La Réunion Island	introduced invasive	Lavergne, Christophe (2006) "Naturalisé"
Mauritius Mautitius Islands (Mauritius and Rodrigues)	Mauritius Island	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (pp. 193-195)

Control: 

Chemical: "A weed in wet open forests and aquatic environments.  Drizzle applications of glyphosate at 1 lb/acre with a good surfactant cleared Wailoa Pond in Hilo (Robert Nishimoto, Division of Aquatic Resources, Hawai‘i Department of Land and Natural Resources) and Hanamā‘ulu Stream on Kaua‘i. A solution of 1-1.5% glyphosate product in water sprayed to wet the foliage provided control on Maui (Hank Oppenheimer, Maui Pine)"  (Motooka et al 2003).

Additional information:  Information from the book "Weeds of Hawaii‘s Pastures and Natural Areas; An Identification and Management Guide" (Motooka et al., 2003). (PDF format).
Photos and additional information at University of Florida, Center for Aquatic and Invasive Plants.
Information from the Purdue University NewCROP web site.

Additional online information about Urochloa mutica is available from the Hawaiian Ecosystems at Risk project (HEAR).

Taxonomic information about Urochloa mutica may be available from the Germplasm Resources Information Network (GRIN).

References:

Bishop Museum (Honolulu). 0. Voucher specimen #BISH 540436 (Florence, J. 8457).

Bishop Museum (Honolulu). 1911. Voucher specimen #BISH 125938 (McGregor, R.C. 489).

Bishop Museum (Honolulu). 1918. Voucher specimen #BISH 125937 (Nelson, P. 293).

Bishop Museum (Honolulu). 1918. Voucher specimen #BISH 125940 (Nelson, P. 331).

Bishop Museum (Honolulu). 1924. Voucher specimen #BISH 700002 (Bryan Jr, E.H. 216).

Bishop Museum (Honolulu). 1924. Voucher specimen #BISH 700006 (Bryan Jr, E.H. 216).

Bishop Museum (Honolulu). 1927. Voucher specimen #BISH 700003 (Gillespie, J.W. 3461).

Bishop Museum (Honolulu). 1927. Voucher specimen #BISH 700005 (Gillespie, J.W. 3163.3).

Bishop Museum (Honolulu). 1932. Voucher specimen #BISH 700004 (Meebold, A. 17097).

Bishop Museum (Honolulu). 1945. Voucher specimen #BISH 700007 (Greenwood, W. 1121).

Bishop Museum (Honolulu). 1946. Voucher specimen #BISH 588986 (Hosaka, E.Y. 2981).

Bishop Museum (Honolulu). 1947. Voucher specimen #BISH 700009 (Smith, A.C. 4749).

Bishop Museum (Honolulu). 1950. Voucher specimen #BISH 588367 (Fosberg, F.R. 31325).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 699999 (Fosberg, F.R. 35276).

Bishop Museum (Honolulu). 1954. Voucher specimen #BISH 31655 (Parham, J.W. 8751).

Bishop Museum (Honolulu). 1954. Voucher specimen #BISH 588985 (Fosberg, F.R. 35576).

Bishop Museum (Honolulu). 1955. Voucher specimen #BISH 31656 (Maya, K. 9457).

Bishop Museum (Honolulu). 1964. Voucher specimen #BISH 125941 (Stone, B.C.M. 5092).

Bishop Museum (Honolulu). 1966. Voucher specimen #BISH 588393 (Evans, M. 1665).

Bishop Museum (Honolulu). 1973. Voucher specimen #BISH 417774 (Whistler, W.A. W341).

Bishop Museum (Honolulu). 1976. Voucher specimen #BISH 410766 (Lamberson, J.; collector's voucher number unknown).

Bishop Museum (Honolulu). 1978. Voucher specimen #BISH 432334 (Stemmermann, R.L. 3220).

Bishop Museum (Honolulu). 1978. Voucher specimen #BISH 432407 (Stemmermann, R.L. 3220).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491185 (Florence, J. 2885).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491711 (Florence, J. 3508).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491726 (Florence, J. 3183).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 492086 (Florence, J. 2835).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 588738 (Fosberg, F.R. 63226).

Bishop Museum (Honolulu). 1983. Voucher specimen #BISH 649487 (Herbst, D.R. 7167).

Bishop Museum (Honolulu). 1983. Voucher specimen #BISH 1000209 (Herbst, D.R. 7167).

Bishop Museum (Honolulu). 1986. Voucher specimen #BISH 572382 (Rinehart, A. LR12289).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 664868 (Sykes, W.R. 474383).

Charles Darwin Research Station. 2005. CDRS Herbarium records.

Edgar, E./Connor, H. 2000. Flora of New Zealand, vol. V: Gramineae. Manaaki Whenua Press. .

Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1987. A geographical checklist of the Micronesian monocotyledonae. Micronesica 20:1-126.

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