|
L., Passifloraceae |
|
Present on Pacific Islands? yes
Primarily a threat at high elevations? no
Risk assessment results: High risk, score: 10 (Go to the risk assessment)
Common name(s): [more details]
|
English: Cuban buttercup, Marilopez, ramgoat dashalong, sage rose, West Indian holly, yellow alder, yellow bells |
|
I-Kiribati: te kaikainea, te kaikainea, te kaikainea, te kaikainea |
|
Marshallese: nõr in jiboñ, nor in jiboñ |
Habit: herb
Description: "Aromatic perennial herbs or shrubs 4-15 dm tall, densely strigose. Leaves clustered toward the tips of the branches, lanceolate to oblong-lanceolate or narrowly elliptic, 4-13 cm long, 2-3 (-5) cm wide, moderately to occasionally sparsely strigillose, margins doubly serrate, petioles 0.8-1.5 cm long, stipules erect, triangular, ca 1.5 mm long. Calyx ca 20 mm long, the tube 5-8 mm long, the lobes lanceolate, apex caudate; petals yellow, sometimes with a brown spot toward base, obovate, 20-35 mm long; ovary ovoid, with 6 small apical tubercles. Capsules ovoid, 7-8 mm long. Seeds narrowly obovoid, slightly curved, ca 2.5 mm long" (Wagner et al., 1999; pp. 1295-1296).
"Herbacée pérenne à base ligneuse ou arbrisseau de 20 cm à 1,5 m de hauteur, reconnaissable à ses petites feuilles à marge dentée regroupées au sommet des tiges et à ses grandes fleurs à 5 pétales de couleur jaune vif" (Meyer, 2008; p. 39).
Habitat/ecology: In Hawaii, "naturalized in dry, disturbed areas, 30-190 m" (Wagner et al., 1999; pp. 1295-1296). In Fiji, "naturalized and locally abundant especially in coconut plantations near sea level" (Smith, 1981; p. 663). In New Caledonia, "envahissant à Thio" (MacKee, 1994; p. 135). In Indonesia, "in sunny dry localities, 1-20 m, especially in the coastal regions: grassy fields, wate places, under coconuts, often on sandy soil, light jungles, as a rule in scattered specimens" (Van Steenis, 1948; pp. 237-238).
Propagation: Seed. Seeds reported to be carried by ants (Staples et al, 2000; p. 30).
Native range: Florida, the West Indies and Tropical America; widely planted as an ornamental and naturalized in the tropics.
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Chile (offshore islands)
Rapa Nui (Easter) Island |
Rapa Nui (Easter) Island (Isla de Pasqua) |
introduced
cultivated |
Meyer, Jean-Yves (2008) (pp. 39, 43)
"Nous l’avons vu également communément plantée comme ornementale dans les jardins de Hanga Roa" (Meyer, 2008; p. 39). |
|
Ecuador (Galápagos Islands)
San Cristóbal Group |
San Cristóbal Island |
introduced
cultivated |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Santa Cruz Group |
Santa Cruz Island |
introduced
cultivated |
Charles Darwin Foundation (2008) |
|
Federated States of Micronesia
Chuuk Islands |
Tol Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (p. 30) |
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (p. 30) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
|
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Waguk, Erick E. (2000) (p. 29) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
invasive cultivated |
Lorence, David H./Flynn, Timothy (2010) (p. 25)
Naturalized at Walung. |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
invasive cultivated |
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 138)
Vouchers cited: M.J. Balick 4250 (NY, PTBG, BISH), F. Sohl 84 (NY, PTBG) Naturalized |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2001) (voucher ID: PTBG 38742)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Kandavu Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663)
Voucher cited: DA 2993 |
|
Fiji
Fiji Islands |
Lakemba Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663)
Voucher cited: Garnock-Jones 961 |
|
Fiji
Fiji Islands |
Mango Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663) |
|
Fiji
Fiji Islands |
Naitamba Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663)
Voucher cited: DA 11799 |
|
Fiji
Fiji Islands |
Naitamba Island |
Bishop Museum (Honolulu) (1956) (voucher ID: BISH 14704)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Ovalau Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663) |
|
Fiji
Fiji Islands |
Taveuni Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663)
Vouchers cited: DA 5714, DA 8904, DA 11514, DA 9109, DA 9623 |
|
Fiji
Fiji Islands |
Taveuni Island |
Bishop Museum (Honolulu) (1954) (voucher ID: BISH 14703)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Taveuni Island |
Bishop Museum (Honolulu) (1958) (voucher ID: BISH 14705)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Vanua Levu Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663)
Vouchers cited: Krauss 1023, Degener & Ordonez 14193, DA 8818, DA 13955 |
|
Fiji
Fiji Islands |
Vanua Levu Island |
Bishop Museum (Honolulu) (1964) (voucher ID: BISH 14702)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Vanua Levu Island |
Bishop Museum (Honolulu) (1941) (voucher ID: BISH 185047)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Vanua Levu Island |
Bishop Museum (Honolulu) (1966) (voucher ID: BISH 185048)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Fiji
Fiji Islands |
Vanua Mbalavu Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663)
Vouchers cited: DA 10220, Garnock-jones 1115 |
|
Fiji
Fiji Islands |
Vanua Mbalavu Island |
Bishop Museum (Honolulu) (1977) (voucher ID: BISH 420632)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
French Polynesia
Marquesas Islands |
Ua Pou (Huapu, Uapou, Uapu) Island |
introduced
cultivated |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
cultivated |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée |
|
French Polynesia
Tuamotu Archipelago |
Raroia Atoll |
introduced
invasive cultivated |
Meyer, Jean-Yves/Fourdrigniez, Marie (2008) (p. 15) |
|
French Polynesia
Austral (Tubuai) Islands |
Tubuai Island |
introduced
cultivated |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée |
|
State of Hawaii
Hawaiian Islands |
Hawaii (Big) Island |
introduced
invasive |
Oppenheimer, Hank L. (2004) (p. 18)
Voucher cited: Oppenheimer H80107 (BISH, PTBG) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1295-1296) |
|
State of Hawaii
Hawaiian Islands |
Maui Island |
introduced
invasive |
Oppenheimer, Hank L. (2004) (p. 18)
West Maui. Vouchers cited: Oppenheimer H50111 (BISH, PTBG), Oppenheimer H50112 (BISH, PTBG), Oppenheimer H60133 (BISH, PTBG), Oppenheimer & Hansen H80301 (BISH, PTBG), Oppenheimer H110110 (BISH, PTBG) |
|
State of Hawaii
Hawaiian Islands |
Molokai Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1295-1296) |
|
State of Hawaii
Hawaiian Islands |
Molokai Island |
introduced
invasive cultivated |
Smith, Albert C. (1981) (p. 663) |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
|
Wester, Lyndon (1992) (p. 151) |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive cultivated |
Frohlich, Danielle/Lau, Alex (2007) (p. 11)
Voucher cited: D. Frohlich s.n. (BISH 725947) |
|
Kiribati
Tungaru (Gilbert) Islands |
Abemama Atoll |
introduced
invasive |
Space, James C./Imada, Clyde T. (2004) (p. 10)
Voucher: Imada 2003-148 (BISH 697760) |
|
Kiribati
Tungaru (Gilbert) Islands |
Abemama Atoll |
Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697760)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Kiribati
Tungaru (Gilbert) Islands |
Butaritari Atoll |
introduced
invasive |
Space, James C./Imada, Clyde T. (2004) (p. 10)
Voucher: Imada 2003-54 (BISH 697669) |
|
Kiribati
Tungaru (Gilbert) Islands |
Butaritari Atoll |
Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697669)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Kiribati
Tungaru (Gilbert) Islands |
Maiana Atoll |
introduced
invasive |
Space, James C./Imada, Clyde T. (2004) (p. 10)
Voucher: Imada 2003-102 (BISH 697717) |
|
Kiribati
Tungaru (Gilbert) Islands |
Maiana Atoll |
Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697717)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Kiribati
Tungaru (Gilbert) Islands |
Tarawa Atoll |
introduced
invasive |
Space, James C./Imada, Clyde T. (2004) (p. 10)
Voucher specimens collected: Imada 2003-25 (BISH 697640); Imada 2003-30 (BISH 697645) |
|
Kiribati
Tungaru (Gilbert) Islands |
Tarawa Atoll |
Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697640)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Kiribati
Tungaru (Gilbert) Islands |
Tarawa Atoll |
Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697645)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Marshall Islands
Ralik Chain |
Kili (Kõle) Atoll |
introduced
invasive cultivated |
Vander Velde, Nancy/Vander Velde, Brian (2006) (p. 12)
"Common; decorative around many houses, some escaping into the wilds". |
|
Marshall Islands
Ralik Chain |
Kwajalein (Kuwajleen) Atoll |
introduced
cultivated |
Whistler, W. A./Steele, O. (1999) (p. 103) |
|
Marshall Islands
Ratak Chain |
Arno Atoll |
introduced
invasive cultivated |
Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 12)
"Common; decorative around houses, some plants becoming ruderal, even escaping into forest areas". |
|
Marshall Islands
Ratak Chain |
Majuro (Mãjro) Atoll |
introduced
invasive cultivated |
Vander Velde, Nancy (2003) (p. 129) |
|
Marshall Islands
Ratak Chain |
Maloelap Atoll | Imada, Clyde (2006) | |
|
Marshall Islands
Ratak Chain |
Mili (Mile) Atoll |
introduced
invasive |
Vander Velde, Nancy/Vander Velde, Brian (2010) (p. A22)
Occasional; near homes, escaping into coconut plantations and ruderal areas, some residents complain that once established, cannot be eradicated, Mili, Bar, Nallo. Voucher 0724201006 (BISH, GUAM, PTBG) |
|
Marshall Islands
Ratak Chain |
Wotje (Wõjjã) Atoll |
introduced
|
Vander Velde, Nancy/Vander Velde, Brian (2009) (p. 3)
Voucher cited: Vander Velde 0516200908 |
|
Marshall Islands
Ratak Chain |
Wotje (Wõjjã) Atoll |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2009) (voucher ID: PTBG 49778)
Taxon name on voucher: Turnera ulmifolia |
|
|
Nauru
Nauru Island |
Nauru Island |
introduced
|
Meyer, Jean-Yves (2000) (p. 99)
"Potential invader". |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Maré |
introduced
invasive cultivated |
MacKee, H. S. (1994) (p. 135)
Voucher cited: Suprin in Veillon 6294 (NOU) |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Tiga |
introduced
invasive cultivated |
MacKee, H. S. (1994) (p. 135)
Voucher cited: MacKee 45911 |
|
New Caledonia
New Caledonia |
New Caledonia Islands |
introduced
|
Meyer, Jean-Yves (2000) (p. 100)
"Potential invader". |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive |
Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 383)
Envahissant à Thio. |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive cultivated |
MacKee, H. S. (1994) (p. 135)
Vouchers cited: MacKee 41650, MacKee 43038 |
|
Palau
Palau (main island group) |
Angaur Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 88) |
|
Palau
Palau (main island group) |
Angaur Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 13) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
invasive cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 87) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
invasive cultivated |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 13) |
|
Palau
Palau (main island group) |
Babeldaob Island |
Bishop Museum (Honolulu) (1996) (voucher ID: BISH 649710)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Palau
Palau (main island group) |
Kayangel (Ngcheangel) Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (pp. 8, 13) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 83) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 13) |
|
Palau
Palau (main island group) |
Malakal Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 83) |
|
Palau
Palau (main island group) |
Malakal Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 13) |
|
Palau
Palau (main island group) |
Ngerkebesang Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 83) |
|
Palau
Palau (main island group) |
Ngerkebesang Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 13) |
|
Palau
Palau (main island group) |
Peleliu Island |
introduced
invasive cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 91) |
|
Palau
Palau (main island group) |
Peleliu Island |
introduced
invasive cultivated |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 13) |
|
Palau
Rock Islands |
Ngerchong Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (pp. 7, 13) |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis (Uvea) Island |
introduced
invasive cultivated |
Meyer, Jean-Yves (2007) (pp. 12, 38)
Cette plante ornementale à grandes fleurs jaune d’or est localement naturalisée en bordure de route et de piste. |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
El Salvador
El Salvador |
El Salvador (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 371) | |
|
Indonesia
Indonesia |
Indonesia (Republic of) |
introduced
invasive |
van Steenis, C. G. G. J. et al (1948) (p. 4:238) |
|
Malaysia
Malaysia |
Malaysia (country of) |
introduced
invasive |
van Steenis, C. G. G. J. et al (1948) (p. 4:238) |
|
Mexico
Mexico |
Mexico (United Mexican States) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Singapore
Singapore |
Singapore (Republic of) |
introduced
invasive |
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 88)
Naturalised |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia (Indian Ocean offshore islands)
Christmas Island Group |
Christmas Island |
introduced
invasive |
Swarbrick, J. T. (1997) (p. 130) |
|
Australia (Indian Ocean offshore islands)
Cocos (Keeling) Islands |
Cocos (Keeling) Islands |
introduced
invasive |
Orchard, Anthony E., ed. (1993) (p. 37) |
|
British Indian Ocean Territory
Chagos Archipelago |
Diego Garcia Island |
introduced
|
Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 154)
Var. elegans Urb. "Recorded by Willis and Gardiner (1931)". |
|
British Indian Ocean Territory
Chagos Archipelago |
Diego Garcia Island |
introduced
invasive cultivated |
Whistler, W. Arthur (1996) (pp. 30, 35) |
|
British Indian Ocean Territory
Chagos Archipelago |
Diego Garcia Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 38066)
Taxon name on voucher: Turnera ulmifolia L. |
|
|
Maldives
Maldive Islands |
Addu Atoll | Spicer, R. A./Newberry, D. MacC. (1979) (p. 12) | |
|
Maldives
Maldive Islands |
Malè Atoll |
Fosberg, F. R. (1957) (p. 27)
"Common generally especially along lagoon shore". Voucher cited: Fosberg 36849 (US) |
|
|
Seychelles
Seychelles Islands |
Agalega Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Stoddart, D. R. (1983) (p. 139)
Vouchers cited: Stoddart 7291 (US), Stoddart 7296 (US) |
|
Seychelles
Seychelles Islands |
Alphonse Island |
Robertson, I. A. D./Robertson, S. A./Fosberg, F. R. (1983) (p. 183)
Voucher cited: Robertson 2854 (K) |
|
|
Seychelles
Seychelles Islands |
Coetivy Island |
Robertson, S. A./Fosberg, F. R. (1983) (p. 155)
Voucher cited: Robertson 3068 (K) |
|
|
Seychelles
Seychelles Islands |
Cousin Island |
introduced
invasive |
Fosberg, F. R. (1983) (p. 29)
Voucher cited: Fosberg 52035 (US) |
|
Seychelles
Seychelles Islands |
D'Arros Island |
Stoddart, D. R./Coe, M. J./Fosberg, F. R. (1979) (p. 23)
Vouchers cited: Gwynne & Wood 927 (EA), Gwynne & Wood 950 (EA), Gwynne & Wood 961 (EA), Stoddart 7213 (US) |
|
|
Seychelles
Seychelles Islands |
Frégate Island |
Robertson, S. A./Todd, D. M. (1983) (p. 61)
Voucher cited: Robertson 2654 |
|
|
Seychelles
Seychelles Islands |
Marie-Louise Island | Wilson, J. R. (1983) (p. 195) | |
|
Seychelles
Seychelles Islands |
Platte Island |
Robertson, S. A./Fosberg, F. R. (1983) (p. 163)
Voucher cited: Robertson 3046 (K) |
|
|
Seychelles
Seychelles Islands |
Poivre Island |
Robertson, S. A./Fosberg, F. R. (1983) (p. 174)
Voucher cited: Robertson 2355 (K) |
|
|
Seychelles
Seychelles Islands |
St. Joseph Island |
Stoddart, D. R./Coe, M. J./Fosberg, F. R. (1979) (p. 46)
Voucher cited: Stoddart 7180 (US) |
|
Comments: See also Turnera subulata.
Additional information:
Additional online information about Turnera ulmifolia is available from the Hawaiian Ecosystems at Risk project (HEAR).
Information about Turnera ulmifolia as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).
Taxonomic information about Turnera ulmifolia may be available from the Germplasm Resources Information Network (GRIN).
References:
Bishop Museum (Honolulu). 1941. Voucher specimen #BISH 185047 (Degener, O. 14193).
Bishop Museum (Honolulu). 1954. Voucher specimen #BISH 14703 (Koroiveibau, D. 8904).
Bishop Museum (Honolulu). 1956. Voucher specimen #BISH 14704 (Morwood, R.B. 11799).
Bishop Museum (Honolulu). 1958. Voucher specimen #BISH 14705 (Parham, J.W. 11514).
Bishop Museum (Honolulu). 1964. Voucher specimen #BISH 14702 (Qoro, I. 13955).
Bishop Museum (Honolulu). 1966. Voucher specimen #BISH 185048 (Krauss, N.L.H. 1023).
Bishop Museum (Honolulu). 1977. Voucher specimen #BISH 420632 (Garnock-Jones, P.J. 307493).
Bishop Museum (Honolulu). 1996. Voucher specimen #BISH 649710 (Herbst, D.R. LR26198).
Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697640 (C.Imada 2003-25).
Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697645 (C.Imada 2003-30).
Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697669 (C.Imada 2003-54).
Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697717 (C.Imada 2003-102).
Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697760 (C.Imada 2003-148).
Charles Darwin Foundation. 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Charles Darwin Foundation, Galapagos, Ecuador.
Charles Darwin Research Station. 2005. CDRS Herbarium records.
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species. Raffles Museum of Biodiversity Research, National University of Singapore. 273 pp.
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