Sphagneticola trilobata (L.) Pruski, Asteraceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Risk assessment results: 

Reject, score: 6 (Go to the risk assessment (Australia))
High risk, score: 13 (Go to the risk assessment (Pacific))

Other Latin names:  Thelechitonia trilobata (L.) H. Rob. & Cautrec.; Wedelia trilobata (L.) Hitchc.

Common name(s): [more details]

Chuukese: atiat

English: creeping ox-eye, Singapore daisy, trailing daisy, wedelia

Kosraean: rosrangrang

Marshallese: ut mõkadkad, ut telia

Palauan: ngesil ra ngebard

Pohnpeian: dihpwoangoahng suwed, ngkahu, tuhke ongohng

Tongan: ‘anselmo

Habit:  herb

Description:  "Creeping, mat-forming perennial herbs; stems rounded, rooting at the nodes, 1-3 (-4) dm long, the flowering portions ascending, coarsely strigose to spreading hirsute, sometimes subglabrous.  Leaves fleshy, usually 4-9 cm long, (1.5-) 2-5 cm wide, irregularly toothed or serrate, usually with a pair of lateral lobes.  Peduncles 3-10 cm long; involucre campanulate-hemispherical, ca. 1 cm high; chaffy bracts lanceolate, rigid; ray florets often 8-13 per head, rays 6-15 mm long; disk corollas 4-5 mm long; pappus a crown of short fimbriate scales.  Achenes tuberculate, 4-5 mm long, few achenes maturing in cultivated plants in Hawai‘i"  (Wagner et al., 1999; pp. 373-374).

Habitat/ecology:  "The plant thrives in open areas with well-drained, moist soil, but it can tolerate dry periods" (Csurhes & Edwards, 1998). Open areas with well-drained soil up to 700 m or more in elevation (up to 1300 m in French Polynesia). A noxious weed in agricultural areas, along roadsides and trails, in open lots, waste places and garbage dumps and other disturbed sites. Also naturalized and invasive along streams, canals, along the borders of mangroves and in coastal strand vegetation. (Thaman, 1999) Forms a dense ground cover, crowding out or preventing regeneration of other species. Cultivated as an ornamental.

"In the wet tropics region of northern Queensland [Australia], the plant is becoming increasingly common along the edges of rainforests, on roadsides and other disturbed areas.  In south-east Queensland and coastal north New South Wales, it has invaded riparian vegetation and disturbed areas of coastal sand dunes."  (Csurhes & Edwards, 1998; p. 201).

Propagation:  Usually vegetatively, but Bill Sykes reports observing mature achenes on plants (pers. com.). Stems form new plants where they touch the ground and pieces readily take root. Commonly spread by dumping of garden waste.

Native range:  "Mexico, Central America, and the islands of the Caribbean; now grown almost worldwide in tropical and other warm places"  (Staples & Herbst, 2005; p. 165).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Manu‘a Islands	Ta‘u Island	introduced invasive cultivated	Space, James C./Flynn, Tim (2000) (p. Field notes.)
American Samoa Tutuila Islands	Tutuila Island	introduced invasive cultivated	Space, James C./Flynn, Tim (2000) (p. 38)
American Samoa Tutuila Islands	Tutuila Island		Bishop Museum (Honolulu) (1972) (voucher ID: BISH 83847) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
American Samoa Tutuila Islands	Tutuila Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1992) (voucher ID: PTBG 48583) Taxon name on voucher: Sphagneticola trilobata
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island	introduced invasive	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane (2000) (p. 7)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 593395) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 593620) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Cook Islands Southern Cook Islands	‘Atiu Island	cultivated	McCormack, Gerald (2011)
Cook Islands Southern Cook Islands	‘Atiu Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. 13)
Cook Islands Southern Cook Islands	Rarotonga Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 13)
Cook Islands Southern Cook Islands	Rarotonga Island		Bishop Museum (Honolulu) (1990) (voucher ID: BISH 665083) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Cook Islands Southern Cook Islands	Rarotonga Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 665339) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Cook Islands Southern Cook Islands	Rarotonga Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 665340) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Federated States of Micronesia Chuuk Islands	Fefan Island	introduced invasive	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (pp. 7, 30)
Federated States of Micronesia Chuuk Islands	Pis Island		Bishop Museum (Honolulu) (1993) (voucher ID: BISH 633457) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Federated States of Micronesia Chuuk Islands	Puluwat (Pwonowót) Atoll	introduced invasive cultivated	Manner, H. I./Mallon, E. (1989) (p. 61) Voucher cited: Manner & Mallon 153 (GUAM)
Federated States of Micronesia Chuuk Islands	Tol Island	introduced invasive	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (pp. 7, 30)
Federated States of Micronesia Chuuk Islands	Weno (Moen) Island	introduced invasive	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (pp. 7, 30)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive	Meyer, Jean-Yves (2000) (p. 90)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive	Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Waguk, Erick E. (2000) (p. 6) Voucher: BMW 5762 (BISH, BRI)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive cultivated	Lorence, David H./Flynn, Timothy (2010) (p. 13) Voucher cited: Lorence 7864 (PTBG) Invasive
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive	Josekutty, P. C./Wakuk, E. E./Joseph, M. J. (2002) (p. 63)
Federated States of Micronesia Kosrae Island	Kosrae Island		Bishop Museum (Honolulu) (2000) (voucher ID: BISH 687682) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Federated States of Micronesia Kosrae Island	Kosrae Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1996) (voucher ID: PTBG 21073) Taxon name on voucher: Sphagneticola trilobata (L.) Hitchc.
Federated States of Micronesia Kosrae Island	Kosrae Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 24672) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive	Meyer, Jean-Yves (2000) (p. 90)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive cultivated	Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 73) Vouchers cited: M.J. Balick 3823 (NY), M.J. Balick 3856 (NY, PTBG), M.J. Balick 4028 (NY, PTBG), T.W. Flynn 6255 (AD, NY, PTBG, US), D.H. Lorence 8057 (MO, NY, PTBG, US) Invasive
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Bishop Museum (Honolulu) (1980) (voucher ID: BISH 505203) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 33554) Taxon name on voucher: Sphagneticola triloba (L.) Pruski
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2026) (voucher ID: PTBG 34871) Taxon name on voucher: Sphagneticola triloba (L.) Pruski
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 24622) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 27787) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 293)
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	introduced invasive	Meyer, Jean-Yves (2000) (p. 90)
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	introduced invasive	Space, James C./Falanruw, Marjorie (1999) (p. Field notes.)
Fiji Fiji Islands	Viti Levu Island	introduced invasive cultivated	Thaman, R. R. (1999) (p. 3)
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Marquesas Islands	Fatu Hiva Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Fatu Hiva Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Vouchers cited: J. Florence 7432 (PAP), J. Florence, D.H. Lorence & W.L. Wagner 9507 (PAP), Adventice
French Polynesia Marquesas Islands	Fatu Hiva Island		Bishop Museum (Honolulu) (1986) (voucher ID: BISH 545498) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Fatu Hiva Island		Bishop Museum (Honolulu) (1988) (voucher ID: BISH 601880) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Fatu Hiva Island		Bishop Museum (Honolulu) (1988) (voucher ID: BISH 1000349) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Fatu Hiva Island		Bishop Museum (Honolulu) (1988) (voucher ID: BISH 1000350) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Fatu Hiva Island		Bishop Museum (Honolulu) (1988) (voucher ID: BISH 1002250) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Marquesas Islands	Mohotani (Mohotane, Mohotane) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive cultivated	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence 4133 (PAP), J. Florence 4193 (PAP) Naturalisée
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 558046) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Tahuata (Tuhuata) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Tahuata (Tuhuata) Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39490) Taxon name on voucher: Sphagneticola trilobata
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: K.R. Wood & S.P. Perlman 6403 (PAP) Naturalisé
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 28584) Taxon name on voucher: Sphagneticola trilobata (L.) Hitchc.
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Society Islands	Huahine Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence & R. Tahuaitu 11711 (PAP) Naturalisée
French Polynesia Society Islands	Huahine Island		Bishop Museum (Honolulu) (1992) (voucher ID: BISH 649993) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Society Islands	Moorea Island	introduced invasive	Welsh, S. L. (1998) (p. 82) Voucher cited: Florence 2126
French Polynesia Society Islands	Moorea Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence 2126 (PAP) Naturalisée
French Polynesia Society Islands	Moorea Island		Bishop Museum (Honolulu) (1981) (voucher ID: BISH 492782) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Welsh, S. L. (1998) (p. 82) Vouchers cited: BRY 24831, BRY 25408
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Vouchers cited: J. Florence 10420 (PAP), J. Florence 3495 (PAP), J. Florence 3681 (PAP) Naturalisée
French Polynesia Society Islands	Raiatea (Havai) Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491380) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Society Islands	Raiatea (Havai) Island		Bishop Museum (Honolulu) (1990) (voucher ID: BISH 644386) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
French Polynesia Society Islands	Taha‘a Island		Fosberg, F. R. (1997) (p. 19)
French Polynesia Society Islands	Tahiti Island		Fosberg, F. R. (1997) (p. 19)
French Polynesia Society Islands	Tahiti Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Society Islands	Tetiaroa Atoll	introduced invasive	Sachet, M.-H./Fosberg, F. R. (1983) (p. 54)
French Polynesia Society Islands	Tupai Island	introduced invasive	Sachet, M.-H. (1983) (p. 24) Voucher cited: Sachet 1967 (US)
French Polynesia Tuamotu Archipelago	Hao (Hau) Atoll	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Tuamotu Archipelago	Moruroa Atoll	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Austral (Tubuai) Islands	Raivavae (Raevavae) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Austral (Tubuai) Islands	Rapa Island	introduced invasive	Welsh, S. L. (1998) (p. 82)
French Polynesia Austral (Tubuai) Islands	Rimatara (Rimitara) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Austral (Tubuai) Islands	Tubuai Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Naturalisée
French Polynesia Austral (Tubuai) Islands	Tubuai Island		Bishop Museum (Honolulu) (1974) (voucher ID: BISH 599630) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 293)
Guam Guam Island	Guam Island	introduced invasive	Meyer, Jean-Yves (2000) (p. 95)
Guam Guam Island	Guam Island	introduced invasive	Fosberg, F. R./Falanruw, M. V. C. (1980) (p. 210) Voucher cited: Fosberg & Moore 58195 (US, BISH)
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1978) (voucher ID: BISH 438141) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
State of Hawaii Hawaiian Islands	Hawai‘i (Big) Island	introduced invasive cultivated	Imada, Clyde T./Staples, George W./Herbst, Derral R. (2000) (p. 10) Voucher cited: C. Imada & T. Cummings 99-14 (BISH)
State of Hawaii Hawaiian Islands	Kaho‘olawe Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)
State of Hawaii Hawaiian Islands	Kaua‘i Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)
State of Hawaii Hawaiian Islands	Lāna‘i Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)
State of Hawaii Hawaiian Islands	Lāna‘i Island	introduced invasive cultivated	Oppenheimer, Hank (2007) (pp. 20-21) Vouchers cited: Oppenheimer H100633 (BISH, PTBG), Oppenheimer H120650 (BISH, PTBG)
State of Hawaii Hawaiian Islands	Maui Island	introduced invasive cultivated	Oppenheimer, Hank L./Bartlett, Randal T. (2002) (p. 4) Vouchers cited: (West Maui) Oppenheimer H99927 (BISH), Oppenheimer H80013 (BISH); (East Maui) Flynn & From 2543 (BISH), Oppenheimer & J. Price H30007 (BISH)
State of Hawaii Hawaiian Islands	Moloka‘i Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)
State of Hawaii Hawaiian Islands	Moloka‘i Island	introduced invasive cultivated	Oppenheimer, Hank (2007) (pp. 20-21) Vouchers cited: Oppenheimer H110606 (BISH, PTBG), Oppenheimer H120616 (BISH, PTBG), Oppenheimer H120630 (BISH, PTBG)
State of Hawaii Hawaiian Islands	Ni‘ihau Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)
Japan (offshore islands) Bonin (Ogasawara) Islands	Bonin (Ogasawara) Islands	introduced	Kato, Hidetoshi (2007)
Kiribati Tungaru (Gilbert) Islands	Butaritari Atoll	introduced invasive cultivated	Space, James C./Imada, Clyde T. (2004) (p. 12) Voucher: Imada 2003-65 (BISH 697680)
Kiribati Tungaru (Gilbert) Islands	Butaritari Atoll		Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697680) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Kiribati Tungaru (Gilbert) Islands	Tarawa Atoll	introduced invasive cultivated	Space, James C./Imada, Clyde T. (2004) (pp. 9, 12) Voucher: Imada 2003-40 (BISH 697655)
Kiribati Tungaru (Gilbert) Islands	Tarawa Atoll		Bishop Museum (Honolulu) (2003) (voucher ID: BISH 697655) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Marshall Islands Ralik Chain	Enewetak (Ãnewetak, Eniwetok) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 293)
Marshall Islands Ralik Chain	Enewetak (Ãnewetak, Eniwetok) Atoll	introduced invasive	Fosberg, F. R./Falanruw, M. V. C. (1980) (p. 210) Voucher cited: Lamberson 29 (US)
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 293)
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll	introduced invasive cultivated	Whistler, W. A./Steele, O. (1999) (p. 39) Vouchers cited: Fosberg 48067, Whistler 11211
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll	introduced invasive	Fosberg, F. R./Falanruw, M. V. C. (1980) (p. 210) Voucher cited: Fosberg 58067 (US)
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1998) (voucher ID: PTBG 43931) Taxon name on voucher: Sphagneticola trilobata
Marshall Islands Ratak Chain	Arno Atoll	introduced invasive	Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 5) "Common; around homes, in former Agriculture station, escaping into ruderal areas ".
Marshall Islands Ratak Chain	Arno Atoll	introduced	Vander Velde, Nancy/Vander Velde, Brian (2009) (p. 4) Voucher cited: Vander Velde 0630200904
Marshall Islands Ratak Chain	Majuro (Mãjro) Atoll	introduced invasive	Vander Velde, Nancy (2003) (p. 72)
Marshall Islands Ratak Chain	Mili (Mile) Atoll	introduced invasive	Vander Velde, Nancy/Vander Velde, Brian (2010) (p. A11) Occasional; near runway, by some residences and other buildings on Mili.
Marshall Islands Ratak Chain	Wotje (Wõjjã) Atoll	introduced	Vander Velde, Nancy/Vander Velde, Brian (2009) (p. 2) Voucher cited: Vander Velde 0515200909
Marshall Islands Ratak Chain	Wotje (Wõjjã) Atoll		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2009) (voucher ID: PTBG 49776) Taxon name on voucher: Sphagneticola trilobata
Nauru Nauru Island	Nauru Island	introduced invasive	Meyer, Jean-Yves (2000) (p. 99)
Nauru Nauru Island	Nauru Island	introduced cultivated	Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 102) Vouchers cited: Fosberg 58609 (US), Hassall 237 (SUVA)
Nauru Nauru Island	Nauru Island	introduced invasive	Fosberg, F. R./Falanruw, M. V. C. (1980) (p. 210) Voucher cited: Fosberg 58609 (US)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive cultivated	MacKee, H. S. (1994) (p. 39) Vouchers cited: MacKee 28336, MacKee 41221
New Caledonia New Caledonia Archipelago	Île Grande Terre		Bishop Museum (Honolulu) (1992) (voucher ID: BISH 655114) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Niue Niue	Niue Island	introduced invasive cultivated	Space, James C./Flynn, Tim (2000) (p. 6)
Niue Niue	Niue Island	introduced invasive	Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 8) Subject of eradication program.
Palau Hatohobei Group	Tobi Island		Bishop Museum (Honolulu) (1992) (voucher ID: BISH 1001361) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Palau Palau (main island group)	Angaur Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 11, 88)
Palau Palau (main island group)	Angaur Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 12)
Palau Palau (main island group)	Babeldaob Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 11, 87)
Palau Palau (main island group)	Babeldaob Island	introduced invasive cultivated	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 12)
Palau Palau (main island group)	Babeldaob Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 610707) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Palau Palau (main island group)	Babeldaob Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 610717) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Palau Palau (main island group)	Babeldaob Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 611191) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Palau Palau (main island group)	Babeldaob Island		Bishop Museum (Honolulu) (1996) (voucher ID: BISH 650941) Taxon name on voucher: Sphagneticola trilobata (L.) Pruski
Palau Palau (main island group)	Babeldaob Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 1000102) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1991) (voucher ID: BNM 9570) Taxon name on voucher: Wedelia trilobata (L.) Hitchc.
Palau Palau (main island group)	Koror Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 11, 83)
Palau Palau (main island group)	Koror Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 12)
Palau Palau (main island group)	Malakal Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 11, 83)
Palau Palau (main island group)	Malakal Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 12)
Palau Palau (main island group)	Ngerkebesang Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 11, 83)
Palau Palau (main island group)	Peleliu Island	introduced invasive cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 11, 91)
Palau Palau (main island group)	Peleliu Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 12)
Samoa Western Samoa Islands	Savai‘i Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 11)
Samoa Western Samoa Islands	Upolu Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 11)
Tokelau Tokelau Islands	Nukunonu Atoll	introduced invasive	? (year unknown) Mike Thorsen, pers. com.
Tonga Tongatapu Group	Tongatapu Island	introduced invasive cultivated	Space, James C./Flynn, Tim (2001) (p. 9)
Tonga Tongatapu Group	Tongatapu Island	introduced invasive cultivated	Thaman, R. R. (1999)
United States (other Pacific offshore islands) Midway Atoll	Midway Atoll	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 373-374, 1864)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia Australia (continental)	New South Wales	introduced invasive cultivated	Csurhes, S./Edwards, R. (1998) (p. 201)
Australia Australia (continental)	Queensland	introduced invasive cultivated	Csurhes, S./Edwards, R. (1998) (p. 201)
China China	Hong Kong	introduced cultivated	Wu, Te-lin (2001) (p. 290)
Colombia Colombia	Colombia (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Costa Rica Costa Rica	Costa Rica (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Ecuador (Mainland) Ecuador	Ecuador (Republic of) (continental)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Guatemala Guatemala	Guatemala (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Honduras Honduras	Honduras (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Japan Japan	Japan (country)	introduced	Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 190) As Wedelia trilobata Hitchc.
Mexico Mexico	Mexico (United Mexican States)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Nicaragua Nicaragua	Nicaragua (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Panama Panama	Panama (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Perú Perú	Perú (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Singapore Singapore	Singapore (Republic of)	introduced invasive	Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 81) Naturalised
Taiwan Taiwan	Taiwan Island	introduced invasive	Ching-I Peng, ed. (2011) "Occasionally naturalized on slopes and along roadsides at lower elevations".

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia (Indian Ocean offshore islands) Christmas Island Group	Christmas Island	introduced invasive cultivated	Swarbrick, J. T. (1997) (p. 131)
French Territory of Mayotte Mayotte Islands	Mayotte Island	introduced invasive	Comité français de l'Union Internationale pour la Conservation de la Nature en France (2010)

Also reported from
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
United States (continental except west coast) United States (other states)	United States (other states)	introduced	U.S. Dept. Agr., Nat. Res. Cons. Serv. (2011) Louisiana
United States (continental except west coast) United States (other states)	USA (Florida)	introduced	U.S. Dept. Agr., Nat. Res. Cons. Serv. (2011)

Comments:  This species was previously widely known as Wedelia trilobata.

Becoming established on Niue (Space & Flynn, 2000). Established at several places on Tongatapu, Tonga (Space & Flynn, 2001). Planted and naturalizing at a number of locations throughout Samoa (Space & Flynn, 2002). Becoming established on Rarotonga and a single cultivated location on ‘Atiu, Cook Islands (Space & Flynn, 2002).

Can cause itching or contact dermatitis if handled and is a hazard to children and pets  (Staples & Herbst, 2005; p. 165).

Control: 

Physical:  "Physical removal can be effective if used repeatedly for an extended period, but is very labor intensive"  (Englberger, 2009; p. 27).

Chemical:  "Sensitive to dicamba and 2,4-D, but probably best at a minimum of 2 lb/acre.  Sensitive to triclopyr in crop oil applied by the drizzle method but probably at 2 lb/acre or more"  (Motooka et al., 2003).  "Herbicides such as triclopyr (Garlon 4) or glyphosate (Roundup) can be used as a foliar spray"  (Englberger, 2009; p. 27).

"Here in Queensland, Australia Wedelia has become a widespread problem plant. A metsulfuron methyl herbicide (600g/kg) has been registered for spraying the weed. The mixing rate is 10g/100L of water plus a surfactant. This gives good control, but it is important to follow-up with touch up treatments to pick up the underground stems which were missed first time round."  (Greg Brown, Queensland Department of Natural Resources, communications to Aliens listserver).

Additional information:
Excerpt from the book "Weeds of Hawaii‘s Pastures and Natural Areas; An Identification and Management Guide" (Motooka et al., 2003). (PDF format).
Information from the Global Invasive Species Database.
Fact sheet from the Government of Queensland, Australia. (PDF format).

Additional online information about Sphagneticola trilobata is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Sphagneticola trilobata as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Sphagneticola trilobata may be available from the Germplasm Resources Information Network (GRIN).

References:

Beleu National Museum (Republic of Palau. Koror.). 1991. Voucher specimen #BNM1991(Herbst, Stemmermann & Canfield 9570).

Bishop Museum (Honolulu). 1972. Voucher specimen #BISH 83847 (Whistler, W.A. W 330).

Bishop Museum (Honolulu). 1974. Voucher specimen #BISH 599630 (Sachet, M.-H. 1967).

Bishop Museum (Honolulu). 1978. Voucher specimen #BISH 438141 (Fosberg, F.R. 58195).

Bishop Museum (Honolulu). 1980. Voucher specimen #BISH 505203 (Fosberg, F.R. 60455).

Bishop Museum (Honolulu). 1981. Voucher specimen #BISH 492782 (Florence, J. 2126).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491380 (Florence, J. 3495).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 558046 (Florence, J. 4133).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 593395 (Herbst, D.R. 6760).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 593620 (Herbst, D.R. 6930).

Bishop Museum (Honolulu). 1986. Voucher specimen #BISH 545498 (Florence, J. 7432).

Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 601880 (Florence 9507).

Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 1000349 (W.L.Wagner 6135).

Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 1000350 (W.L.Wagner 6172).

Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 1002250 (Florence 9507).

Bishop Museum (Honolulu). 1990. Voucher specimen #BISH 644386 (Florence 10420).

Bishop Museum (Honolulu). 1990. Voucher specimen #BISH 665083 (Sykes, W.R. 467850).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 610707 (Herbst, D.R. 9570).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 610717 (Herbst, D.R. 9511).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 611191 (Herbst, D.R. 9570).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 665339 (Sykes, W.R. 474388).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 665340 (Sykes, W.R. 474402).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 1000102 (Herbst, D.R. 9511).

Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 649993 (Florence 11711).

Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 655114 (Koyama, H. 8228).

Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 1001361 (A.K.Kepler 92-49).

Bishop Museum (Honolulu). 1993. Voucher specimen #BISH 633457 (A.K.Kepler 93-56).

Bishop Museum (Honolulu). 1996. Voucher specimen #BISH 650941 (Singeo, Y. LR26656).

Bishop Museum (Honolulu). 2000. Voucher specimen #BISH 687682 (Waterhouse, B.M. BMW5762).

Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697655 (C.Imada 2003-40).

Bishop Museum (Honolulu). 2003. Voucher specimen #BISH 697680 (C.Imada 2003-65).

Ching-I Peng, ed. 2011. Digital flora of Taiwan (online resource).

Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species. Raffles Museum of Biodiversity Research, National University of Singapore. 273 pp.

Comité français de l'Union Internationale pour la Conservation de la Nature en France. 2010. Les espéces envahissantes en outre-mer (online resource).

Csurhes, S./Edwards, R. 1998. Potential environmental weeds in Australia: Candidate species for preventative control. Canberra, Australia. Biodiversity Group, Environment Australia. 208 pp.

Englberger, Konrad. 2009. Invasive weeds of Pohnpei: A guide for identification and public awareness. Conservation Society of Pohnpei. 29 pp.

Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. 2011. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP). (online resource).

Fosberg, F. R. 1997. Preliminary checklist of the flowering plants and ferns of the Society Islands. Ed. by David R. Stoddart. U. Cal. Berkeley.

Fosberg, F. R./Falanruw, M. V. C. 1980. Noteworthy Micronesian plants 4. Micronesica 16(2):201-210.

Fosberg, F. R./Otobed, D./Sachet, M.-H./Oliver, R. L./Powell, D. A./Canfield, J. E. 1980. Vascular plants of Palau with vernacular names. Department of Botany, The Smithsonian Institution, Washington, D.C. 43 pp.

Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.

Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. 2010. Checklist of the vascular plants of Pohnpei with local names and uses. Allertonia, in press. National Tropical Botanical Garden, Lawai, Hawai‘i. 146 pp.

Imada, Clyde T./Staples, George W./Herbst, Derral R. 2000. New Hawaiian plant records for 1999. In: Evenhuis, Neal L. and Eldredge, Lucius G., eds. Records of the Hawaii Biological Survey for 1999. Part 1: Articles. Bishop Museum Occasional Papers. 63:9-16.

Josekutty, P. C./Wakuk, E. E./Joseph, M. J. 2002. Invasive/weedy angiosperms in Kosrae, Federated States of Micronesia. In: Invasive species and their management. Micronesica Suppl. 6:61-65.

Kato, Hidetoshi. 2007. Herbarium records of Makino Herbarium, Tokyo Metropolitan University. Personal communication.

Lorence, David H./Flynn, Timothy. 2010. Checklist of the plants of Kosrae. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawai‘i. 26 pp.

Lorence, David H./Wagner, Warren L. 2008. Flora of the Marquesas Islands. National Tropical Botanical Garden and the Smithsonian Institution. Online database.

MacKee, H. S. 1994. Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie. Muséum National d'Histoire Naturelle, Paris, 164 p.

Manner, H. I./Mallon, E. 1989. An Annotated list of the vascular plants of Pulawat Atoll. Micronesica 22:23-63.

McCormack, Gerald. 2011. Cook Islands Biodiversity Database, Version 2007.2. Cook Islands Natural Heritage Trust, Rarotonga.

Meyer, Jean-Yves. 2000. Preliminary review of the invasive plants in the Pacific islands (SPREP Member Countries). In: Sherley, G. (tech. ed.). Invasive species in the Pacific: A technical review and draft regional strategy. South Pacific Regional Environment Programme, Samoa. 190 pp.

Mito, Toshikazu/Uesugi, Tetsuro. 2004. Invasive alien species in Japan: the status quo and the new regulation for prevention of their adverse effects. Global Environmental Research 8(2)/2004: 171-191.

Motooka, Philip/Castro, Luisa/Nelson, Duane/Nagai, Guy/Ching, Lincoln. 2003. Weeds of Hawaii‘s Pastures and Natural Areas; An Identification and Management Guide. College of Tropical Agriculture and Human Resources, University of Hawaii at Manoa. 184 pp.

National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.). 1997. Voucher specimen #PTBG33554(M. J. Balick 3856).

National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.). 2026. Voucher specimen #PTBG34871(M. J. Balick 4028).

Oppenheimer, Hank. 2007. New plant records from Moloka‘i, Lāna‘i, Maui, and Hawai‘i for 2006. In: Evenhuis, Neal L. and Eldredge, Lucias G., eds. Records of the Hawaii Biological Survey for 2006. Part 2: Notes. Bishop Museum Occasional Papers 96:17-34.

Oppenheimer, Hank L./Bartlett, Randal T. 2002. New plant records from the main Hawaiian Islands. In: Evenhuis, Neal L. and Eldredge, Lucius G., eds. Records of the Hawaii Biological Survey for 2000. Part 2: Notes. Bishop Museum Occasional Papers. 69:1-14.

Sachet, M.-H. 1983. Botanique de l'ile de Tupai, Iles de la Societe. Atoll Research Bulletin No. 276. Smithsonian Institution, Washington. 26 pp. + photos.

Sachet, M.-H./Fosberg, F. R. 1983. An ecological reconnaissance of Tetiaroa Atoll, Society Islands. Atoll Research Bulletin No. 275. Smithsonian Institution, Washington. 67 pp. + photos.

Santos, Ioanis. 1998. Wedelia trilobata on Pohnpei. Aliens 7:3.

Space, James C./Falanruw, Marjorie. 1999. Observations on invasive plant species in Micronesia. USDA Forest Service, Honolulu. Report to the Pacific Islands Committee, Council of Western State Foresters. USDA Forest Service, Honolulu. 32 pp.

Space, James C./Flynn, Tim. 2000. Observations on invasive plant species in American Samoa.  USDA Forest Service, Honolulu. 51 pp.

Space, James C./Flynn, Tim. 2000. Report to the Government of Niue on invasive plant species of environmental concern.  USDA Forest Service, Honolulu. 34 pp.

Space, James C./Flynn, Tim. 2001. Report to the Kingdom of Tonga on invasive plant species of environmental concern.  USDA Forest Service, Honolulu. 78 pp.

Space, James C./Flynn, Tim. 2002. Report to the Government of the Cook Islands on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 146 pp.

Space, James C./Flynn, Tim. 2002. Report to the Government of Samoa on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 83 pp.

Space, James C./Imada, Clyde T. 2004. Report to the Republic of Kiribati on invasive plant species on the islands of Tarawa, Abemama, Butaritari and Maiana. Cont. no. 2003-006 to the Pac. Biol. Surv. USDA Forest Service and Bishop Museum, Honolulu. 103 pp.

Space, James C./Lorence, David H./LaRosa, Anne Marie. 2009. Report to the Republic of Palau: 2008 update on Invasive Plant Species. USDA Forest Service, Hilo, Hawai‘i. 227 pp.

Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate. 2004. Report to the Government of Niue and the United Nations Development Programme: Invasive plant species on Niue following Cyclone Heta. UNDP NIU/98/G31 - Niue Enabling Activity. 80 pp.

Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane. 2000. Invasive plant species on Rota, Commonwealth of the Northern Mariana Islands.  USDA Forest Service, Honolulu. 31 pp.

Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. 2000. Invasive plant species in Chuuk, Federated States of Micronesia.  USDA Forest Service, Honolulu. 41 pp.

Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Waguk, Erick E. 2000. Invasive plant species on Kosrae, Federated States of Micronesia.  USDA Forest Service, Honolulu. 43 pp.

Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar. 2003. Report to the Republic of Palau on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 174 pp.

Staples, George W./Herbst, Derral R. 2005. A tropical garden flora: plants cultivated in the Hawaiian Islands and other tropical places. Bishop Museum Press, Honolulu. 908 pp.

Swarbrick, J. T. 1997. Environmental weeds and exotic plants on Christmas Island, Indian Ocean: a report to Parks Australia. 101 pp. plus appendix.

Swarbrick, John T. 1997. Weeds of the Pacific Islands. Technical paper no. 209. South Pacific Commission, Noumea, New Caledonia. 124 pp.

Thaman, R. R. 1999. Wedelia trilobata: Daisy invader of the Pacific Islands. IAS Technical Report 99/2. Institute of Applied Science, University of the South Pacific, Suva, Fiji. 12 pp.

Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. 1994. The flora of Nauru. Smithsonian Institution, Washington. Atoll Research Bulletin 392:1-223.

U.S. Dept. Agr., Agr. Res. Serv. 2011. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.

U.S. Dept. Agr., Nat. Res. Cons. Serv. 2011. The PLANTS Database. National Plant Data Center, Baton Rouge, LA 70874-4490 USA.

Vander Velde, Nancy. 2003. The vascular plants of Majuro Atoll, Republic of the Marshall Islands. Smithsonian Institution, Atoll Research Bulletin No. 503:1-141.

Vander Velde, Nancy/Vander Velde, Brian. 2007. Arno plants [preliminary working listing of plant species]. Unpublished checklist, 13 pp.

Vander Velde, Nancy/Vander Velde, Brian. 2009. Specimens from Likiep, Majuro, Wotje and Arno (unpublished checklist). 4 pp.

Vander Velde, Nancy/Vander Velde, Brian. 2010. Survey of Invasive Plant Species and Other Plants of Mili Atoll, Marshall Islands. USDA Forest Resource Management Grant Program. Unpublished report. 6 pp, plus tabulation of plants of Mili Atoll, 23 pp.

Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawai‘i Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).

Welsh, S. L. 1998. Flora Societensis: A summary revision of the flowering plants of the Society Islands. E.P.S. Inc., Orem, Utah. 420 pp.

Whistler, W. A. 1991. The ethnobotany of Tonga: The plants, their Tongan names, and their uses. Bishop Museum Bulletin in Botany 2. Bishop Museum Press, Honolulu. 155 pp. ISBN 0-930897-57-9.

Whistler, W. A./Steele, O. 1999. Botanical survey of the United States of America Kwajalein Atoll (USAKA) Islands. Prepared for Oak Ridge Institute for Science and Education and the U. S. Army Environmental Center. 111 pp.

Wu, Te-lin. 2001. Check List of Hong Kong Plants. Hong Kong Herbarium and the South China Institute of Botany. Agriculture, Fisheries and Conservation Department Bulletin 1 (revised). 384 pp.