Sida acuta Burm.f., Malvaceae

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Present on Pacific Islands?  yes

Threat only at high elevations?  no

Risk assessment results:  Reject, score: 19 (Go to the risk assessment)

Common name(s): [more details]

Austral: brum, purumu, purumu, purumu

Bicolano: surusighid

Chamorro: escobilla, escobilla adumelon, escobilla papago, escobilla papagu

Chuukese: sioinen, siyóyinen

English: broom grass, broom weed, clock plant, Paddy's lucerne, southern sida, spiny-head sida

Fijian: deni vuaka, ndeni vuaka, ndeniosa, qawenilawe

Filipino: escuba

French: herbe à balais, herbe à paniers

Ilocano: basbasot

Maori (Cook Islands): pulumi, purūmu

Marquesan: au hopa, hopa, hopa, hopa, kopa, puehu, puehu enana

Niuean: mautofu, motofu

Palauan: keak

Pohnpeian: kahwinyotz

Samoan: mautofu

Spanish: babosilla, escoba, escoba blanca, escoba negra, escobilla, escobilla blanca, escobilla cabezuda, escobilla negra, huinar, malva colorado, malva de castilla

Tagalog: walis-walisan

Tahitian: burume, porumu

Tongan: te‘ehoosi, te‘ekosi

Habit:  herb

Description:  "A small, erect, much branched, perennial shrub or herb; ranging from 30 to 100 cm in height, with a strong taproot; stem and branches flattened at the extremities, fibrous, almost woody at times; leaves alternate, slender, lanceolate, acute, margins toothed, 1.2 to 9 cm or more long, 0.5 to 4 cm wide, lower surface smooth or with sparse, short, branched, starlike (stellate) hairs, with fairly prominent veins; petiole 3 to 6 mm long, hairy, with a pair of stipules, at least one lanceolate-linear, 1 to 2 mm broad, three- to six-nerved, often curved, finely hairy, the other stipule narrower, one- to four-nerved; flowers 1 to 2 cm in diameter, solitary or in densely crowded axillary heads; pedicels 3 to 8 mm, slender, jointed near the middle; calyx five-lobed; sepals pale green, triangular, acute, about 6 mm long; petals five, joined at base, 6 to 9 mm or more long, light yellow, yellow, or pale orange, with a shallow  notch at the apex; ovary eight-celled, one-ovulate; stamens of many filaments arising from a tube; style divided into six branches; fruit a capsule, 3 to 4.5 mm in diameter, rough, consisting  of five to eight (rarely more) carpels which break at maturity into equal, one-seeded, segments (mericarp), each of which has two glabrous or nearly glabrous awns or beaks 1 to 1.5 mm long; seeds small, roughly triangular, 1.5 mm long, with a deep depression on each of the sides, reddish brown or black.  Sida acuta is often confused with Sida rhombifolia, but can be distinguished from that species by its smaller and narrower lanceolate leaves (those of S. rhombifolia are rhomboid or diamond shaped), the shorter pedicels of the flower, and by the two awns of the carpels (S. rhombifolia has one awn or none)" (Holm et al., 1977; p. 426).

"A low shrub to 1 m tall (rarely 2 m) with slender tough stems; leaves lanceolate or linear-lanceolate, acute, obtuse or rounded at base, 1.5-6 cm long; 5-20 mm wide, sharply serrate, stellate-pubescent beneath; petioles 3-5 mm long; blade 3-nereved from base, pinnately branched above the base; stipules linear-acute, about 1 cm long; flowers 1 or 2 in axils, pedicels to about 1 cm long; corolla about 6-14 mm broad, yellow; fruit 4-5 mm broad, 6-8-carpellate; seeds dark reddish-brown, 1.5 mm long; ripe carpels with 2 beaks, grooved on the back"  (Stone, 1970; p. 415).

Habitat/ecology:  "Mainly a weed of the tropics and subtropics.  Although it grows widely throughout the tropics, it can be found at higher elevations.  It has been reported up to 1,500 m in Indonesia.  The weed is frequently found in pastures, wastelands, cultivated lands, roadsides, lawns, and in planted forests.  Once the plant becomes established, it is very competitive, holding and denying sites to other plants. It does appear to do best in disturbed habitats.  The plant grows well in many soils, including some heavy clays, and can tolerate dry as well as high rainfall conditions.  In Fiji it is most aggressive in the dry zones.  It has a deep taproot which can withstand drought, mowing, or shallow infrequent tillage.  Tough woody stems also aid in its persistence" (Holm et al., 1977; p. 426).  "A major weed of pastures and roadsides and also important in orchards and plantations, though not in cultivation" (Swarbrick, 1997; p. 18).  "Occasional in waste areas and roadsides" (Whistler, 1988; p. 29).

In Guam, "common around villages in waste ground"  (Stone, 1970; p. 415).  In Hawai‘i, "naturalized in open and shaded sites, from near sea level up to ca. 790 m" (Wagner et al., 1999; p. 897).  In Fiji, "naturalized in waste places, plantations, canefields, and pastures and along roadsides" (Smith, 1981; p. pp. 436-437). In Tonga, "a rather common waste-area weed" (Yuncker, 1959; p. 181).

Propagation:  Seed.  "The awned mericarps are spread by adhering to clothing and livestock, in mud on vehicles, and as contaminants in hay and seed crops" (Smith, 2002; pp. 40-41).

Native range:  Unknown; a pantropical weed, perhaps indigenous in Central America (Smith, 1981; pp. 436-437).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Tutuila Islands	Tutuila Island		Swarbrick, John T. (1997) (p. 96)
American Samoa Tutuila Islands	Tutuila Island		Anon. (1988) (p. 3)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Agrigan (Agrihan) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164) S. acuta x rhombifolia?
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Alamagan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Alamagan Island	introduced invasive	Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 33) Vouchers cited: Falanruw 1941 (US), Fosberg 31702 (US), Fosberg 31719 (US)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Alamagan Island	introduced	Ohba, T. (1994) (p. 60) Voucher cited: CBM-BS-58865
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Anatahan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Anatahan Island	introduced invasive	Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 33) Voucher cied: Falanruw 1647
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Pagan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Pagan Island	introduced invasive	Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 33) Vouchers cited: Moore 408 (US), Fosberg 31362 (US), Anderson 563 (US)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Sarigan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Sarigan Island	introduced invasive	Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 33) Vouchers cited: Falanruw 1724 (US), Evans 2433 (US, BISH, Fo, NY)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Tinian Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Cook Islands Southern Cook Islands	Ma‘uke Island		McCormack, Gerald (2007)
Cook Islands Southern Cook Islands	Miti‘aro Island		McCormack, Gerald (2007)
Ecuador (Galápagos Islands) Floreana Group	Floreana Island	introduced invasive	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Isabela Group	Volcan Alcedo	introduced invasive	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Isabela Group	Volcan Cerro Azul	introduced invasive	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Isabela Group	Volcan Sierra Negra	introduced invasive	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Santa Cruz Group	Santa Cruz Island	introduced invasive	Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands) Santiago Group	Santiago Island	introduced invasive	Charles Darwin Research Station (2005)
Federated States of Micronesia Chuuk Islands	Dublon (Tonowas) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Federated States of Micronesia Chuuk Islands	Puluwat (Pwonowót) Atoll	introduced invasive	Manner, H. I./Mallon, E. (1989) (p. 52) Voucher cited: Manner & Mallon 267 (GUAM)
Federated States of Micronesia Chuuk Islands	Tol Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Federated States of Micronesia Chuuk Islands	Weno (Moen) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive	Lorence, David H./Flynn, Tim (2005) (p. 17) Voucher cited: Lorence 7865 (PTBG)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Glassman, S. F. (1952) (p. 68) Voucher cited: Glassman 2789 (US)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Lorence, David H./Flynn, Tim (1998) (p. 24)
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Fiji Fiji Islands	Ovalau Island	introduced invasive	Smith, Albert C. (1981) (pp. 436-437) Vouchers cited: US 12252, DA 1135
Fiji Fiji Islands	Taveuni Island	introduced invasive	Smith, Albert C. (1981) (pp. 436-437) Vouchers cited: Gillespie 4795.5, DA 5744
Fiji Fiji Islands	Vanua Levu Island	introduced invasive	Smith, Albert C. (1981) (pp. 436-437) Voucher cited: Greenwood 674
Fiji Fiji Islands	Vanua Mbalavu Island	introduced invasive	Smith, Albert C. (1981) (pp. 436-437) Voucher cited: Garnock-Jones 990
Fiji Fiji Islands	Viti Levu Island	introduced invasive	Smith, Albert C. (1981) (pp. 436-437) Vouchers cited: Greenwood 99, DA 2914, DA 10435, DA 2810, DA 2908, DA 9145, DA 10739, DA 3030, DA 9940
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Marquesas Islands	Eiao Island	introduced invasive	Florence, J. (2004) (pp. 235-238)
French Polynesia Marquesas Islands	Fatu Hiva Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Marquesas Islands	Hatuta'a Island	introduced invasive	Florence, J. (2004) (pp. 235-238)
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive	Florence, J. (2004) (pp. 235-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Florence, J. (2004) (pp. 235-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Marquesas Islands	Tahuata (Tuhuata) Island	introduced invasive	Florence, J. (2004) (pp. 235-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Florence, J. (2004) (pp. 235-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive	Florence, J. (2004) (pp. 235-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Bora Bora Island	introduced invasive	Florence, J. (2004) (pp. 235-238)
French Polynesia Society Islands	Huahine Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Maupiti (Maurua) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Maupiti (Maurua) Island	introduced invasive	Fosberg, F. R./Sachet, M.-H. (1987) (p. 48) Voucher cited: Fosberg 64803 (US) Rare
French Polynesia Society Islands	Mehetia (Mahetia, Meetia) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Moorea Island	introduced invasive	Florence, J. (2004) (pp. 235-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Taha‘a Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Tahiti Island	introduced invasive	Welsh, S. L. (1998) (p. 178) Voucher cited: Quayle 109
French Polynesia Society Islands	Tahiti Island	introduced invasive	Florence, J. (2004) (pp. 238-240) also subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Society Islands	Tupai (Tubai) Island	introduced invasive	Florence, J. (2004) (pp. 238-240)
French Polynesia Society Islands	Tupai (Tubai) Island	introduced	Sachet, M.-H. (1983) (p. 20) Voucher cited: Grant 4848 (BISH)
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) ?
French Polynesia Tuamotu Archipelago	Niau Atoll	introduced invasive	Florence, J. (2004) (pp. 238-240)
French Polynesia Tubuai (Austral) Islands	Raivavae (Raevavae) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Tubuai (Austral) Islands	Rapa Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Tubuai (Austral) Islands	Rurutu Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
French Polynesia Tubuai (Austral) Islands	Tubuai (Tupuai) Island	introduced invasive	Florence, J. (2004) (pp. 238-240) subsp. carpinifolia J. Borssum-Wallkes
Guam Guam Island	Guam Island	introduced invasive	Stone, Benjamin C. (1970) (p. 415) Vouchers cited: Stone 4142 (GUAM), Stone 4995 (GUAM)
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
State of Hawaii Hawaiian Islands	Hawai‘i (Big) Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 897)
State of Hawaii Hawaiian Islands	Kaua‘i Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 897) Voucher cited: Heller 2424 (BISH,CU)
State of Hawaii Hawaiian Islands	Maui Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 897)
State of Hawaii Hawaiian Islands	Moloka‘i Island	introduced	Wester, Lyndon (1992) (p. 142)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 897)
Japan (offshore islands) Bonin (Ogasawara) Islands	Bonin (Ogasawara) Islands		Kato, Hidetoshi (2007)
Kiribati Banaba Island	Banaba (Ocean) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Kiribati Tungaru (Gilbert) Islands	Maiana Atoll	introduced invasive	Space, James C./Imada, Clyde T. (2004) (p. 13) Voucher specimen collected: Imada 2003-111 (BISH 697722)
Marshall Islands Ralik Chain	Rongelap (Roñlap) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Marshall Islands Ralik Chain	Ujelang Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Marshall Islands Ralik Chain	Ujelang Atoll	introduced invasive	Fosberg, F. R. (1955) (p. 15) Voucher cited: Fosberg 34180 (US)
Nauru Nauru Island	Nauru Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164) also S. acuta x rhombifolia?
Nauru Nauru Island	Nauru Island	introduced invasive	Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 167) Vouchers cited: Fosberg 58615 (US), Fosberg 58649 (US), Fosberg 58806 (US), Thaman & Manner 23 (SUVA), Thaman & Manner 96 (SUVA), Thaman & Manner 22303 (SUVA), Swarbrick 9558 (BRIU), Swarbrick 9582 (BRIU)
New Caledonia Îles Bélep (Belep Islands)	Île Art	introduced	MacKee, H. S. (1994) (p. 98) Voucher cited: Montrouzier 8 (LY)
New Caledonia Îles Loyauté (Loyalte Islands)	Îles Loyauté (Loyalty Islands)	introduced invasive	Tassin, Jacques (2005)
New Caledonia Îles Loyauté (Loyalte Islands)	Îles Ouvéa (Ouvea Atoll)	introduced	MacKee, H. S. (1994) (p. 98) Voucher cited: Schmid 2338 (NOU)
New Caledonia Îles Loyauté (Loyalte Islands)	Île Tiga	introduced	MacKee, H. S. (1994) (p. 98) Voucher cited: MacKee 45851
New Caledonia New Caledonia	New Caledonia Islands	introduced invasive	Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 382) Rudérale, nombreuses localités
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	Tassin, Jacques (2005)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced	MacKee, H. S. (1994) (p. 98) Vouchers cited: Pancher s.n., Vieillard 139, MacKee 4055, Toutain 3463 (NOU)
Niue Niue	Niue Island	introduced invasive	Sykes, W. R. (1970) (pp. 116-117) Vouchers cited: CHR 170352, CHR 170345
Niue Niue	Niue Island	introduced invasive	Whistler, W. A. (1988) (p. 29)
Niue Niue	Niue Island	introduced invasive	Yuncker, T. G. (1943) (p. 81) Voucher cited: Yuncker 10150 (BISH) "Occasional along roadsides, in waste areas and plantations".
Niue Niue	Niue Island	introduced invasive	Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 47)
Palau Palau (main island group)	Angaur Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (pp. 163-164)
Palau Palau (main island group)	Angaur Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 104)
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 102)
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)		Waterhouse, D. F. (1997) (p. 65)
Philippines Philippine Islands	Philippine Islands		Waterhouse, D. F. (1993) (pp. 68, 78)
Samoa Western Samoa Islands	Western Samoa Islands		Whistler, W. A. (1988) (p. 29)
Solomon Islands Solomon Islands	Solomon Islands		Swarbrick, John T. (1997) (p. 96)
Solomon Islands Solomon Islands	Solomon Islands	introduced invasive	Hancock, I. R./Henderson, C. P. (1988) (p. 109)
Tonga Ha‘apai Group	Ha‘apai Islands		Whistler, W. A. (1992) (p. 42) Voucher cited: Soakai 571
Tonga Ha‘apai Group	Lifuka Island		Yuncker, T. G. (1959) (p. 181) Voucher cited: Yuncker 15977
Tonga Ha‘apai Group	Nomuka Island		Yuncker, T. G. (1959) (p. 181) Voucher cited: Yuncker 15802
Tonga Tonga Islands	Tonga Islands		Yuncker, T. G. (1959) (p. 181) "Throughout Tonga".
Tonga Vava‘u Group	Vava‘u Island		Whistler, W. A. (1992) (p. 42) Voucher cited: Croby 3
Tonga Vava‘u Group	Vava‘u Island		Yuncker, T. G. (1959) (p. 181) Cited: Crosby.
Vanuatu New Hebrides Islands	Vanuatu (Republic of)		Swarbrick, John T. (1997) (p. 96)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia Australia (continental)	Northern Territory	introduced invasive	Smith, Nicholas M. (2002) (pp. 40-41)
Australia Australia (continental)	Queensland	introduced invasive	Smith, Nicholas M. (2002) (pp. 40-41)
Cambodia Cambodia	Cambodia (Kingdom of)		Waterhouse, D. F. (1993) (pp. 68, 78)
China China	China (People's Republic of)	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (p. 428)
Colombia Colombia	Colombia (Republic of)		Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (pp. 334-335)
El Salvador El Salvador	El Salvador (Republic of)		Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (p. 429)
Honduras Honduras	Honduras (Republic of)		Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (pp. 334-335)
Indonesia Indonesia	Indonesia (Republic of)		Waterhouse, D. F. (1993) (pp. 68, 78)
Malaysia Malaysia	Malaysia (country of)		Waterhouse, D. F. (1993) (pp. 68, 78)
Mexico Mexico	Mexico (United Mexican States)		Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (p. 428)
Panama Panama	Panama (Republic of)		Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (pp. 334-335)
Perú Perú	Perú (Republic of)		Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (p. 428)
Singapore Singapore	Singapore (Republic of)		Waterhouse, D. F. (1993) (pp. 68, 78)
Taiwan Taiwan	Taiwan Island	introduced invasive	Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. (1977) (p. 428)
Thailand Thailand	Thailand (Kingdom of)		Waterhouse, D. F. (1993) (pp. 68, 78)
Vietnam Vietnam	Vietnam (Socialist Republic of)		Waterhouse, D. F. (1993) (pp. 68, 78)

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia (Indian Ocean offshore islands) Christmas Island Group	Christmas Island	native	Orchard, Anthony E., ed. (1993) (p. 18)
Australia (Indian Ocean offshore islands) Cocos (Keeling) Islands	Cocos (Keeling) Islands	native	Orchard, Anthony E., ed. (1993) (p. 38) North Keeling Island
British Indian Ocean Territory Chagos Archipelago	Diego Garcia Island	introduced	Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 154) Vouchers cited: Stoddart 863 (K, US), Rhyne 776 (US)
Mauritius Mautitius Islands (Mauritius and Rodrigues)	Mauritius Island		Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (pp. 334-335)

Comments:  A declared noxious weed in Western Australia and the Northern Territory (Smith, 2002; pp. 40-41).

Control: 

Physical: "Controlled by chipping or cultivation. Suppressed by slashing and vigorous, well-managed, competitive pasture species. Tolerates some shade" (Swarbrick, 1997; p. 29).

Chemical: "Susceptible to: 1) residual herbicides, including 2,4-D, glyphosate, dicamba and fluroxypyr at standard rates; 2) translocated herbicides, including 2,4-D, glyphosate, dicamba and fluroxypyr at standard rates" (Swarbrick, 1997; p. 29).

Biological: "Possible natural enemies are discussed by Waterhouse and Norris (1987). Work is in progress in Australia using the leaf-eating beetle, Calligrapha pantherium" (Swarbrick, 1997; p. 29).

Additional information:  Additional online information about Sida acuta is available from the Hawaiian Ecosystems at Risk project (HEAR).

Taxonomic information about Sida acuta may be available from the Germplasm Resources Information Network (GRIN).

References:

Anon. 1988. Weed list for American Samoa by cropping system. Checklist dated 07/13/88, origin unknown. 3 pp.

Charles Darwin Research Station. 2005. CDRS Herbarium records.

Falanruw, Marjorie C./Maka, Jean E./Cole, Thomas G./Whitesell, Craig D. 1990. Common and scientific names of trees and shrubs of Mariana, Caroline, and Marshall Islands. Pac. SW Forest and Range Expt. Stn. Resource Bulletin PSW-67. 91 pp.

Florence, J. 2004. Flore de la Polynésie française, Vol. 2. Paris. IRE Editions, Publications Scientifiques, Collection Faune et Flore Tropicales 41. 503 pp.

Fosberg, F. R. 1955. Northern Marshall Islands Expedition, 1951-1952. Land biota: vascular plants. Atoll Research Bulletin No. 39. Pacific Science Board, National Academy of Sciences, Washington. 22 pp.

Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. 1975. Vascular flora of the northern Marianas Islands. Smithsonian Contrib. Bot. 22:1-45.

Fosberg, F. R./Sachet, M.-H. 1987. Flora of Maupiti, Society Islands. The Smithsonian Institution. Atoll Research Bulletin 294:1-70.

Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.

Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe. 1996. Conséquences des introductions d'espèces animales et végétales sur la biodiversité en Nouvelle-Calédonie. Rev. Ecol. (Terre Vie) 51:375-401.

Glassman, S. F. 1952. The flora of Ponape. Bishop Mus. Bull. 209: 1-152.

Hancock, I. R./Henderson, C. P. 1988. Flora of the Solomon Islands. Research Bulletin No. 7. Ministry of Agriculture and Lands, Honiara. 203 pp.

Henty, E. E./Pritchard, G. H. 1975. Weeds of New Guinea and their control. 2nd edition. Department of Forests, Division of Botany, Botany Bull. No. 7. Lae, Papua New Guinea. 180 pp.

Holm, Leroy G./Plucknett, D. L./Pancho, J. V./Herberger, J. P. 1977. The world’s worst weeds: distribution and biology. East-West Center/University Press of Hawaii. 609 pp.

Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. 1979. A geographical atlas of world weeds. John Wiley & Sons, New York. 391 pp.

Kato, Hidetoshi. 2007. Herbarium records of Makino Herbarium, Tokyo Metropolitan University. Personal communication.

Lorence, David H./Flynn, Tim. 1998. Checklist of the plants of Pohnpei. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawai‘i. 21 pp.

Lorence, David H./Flynn, Tim. 2005. Checklist of the plants of Kosrae. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawai‘i. 23 pp.

MacKee, H. S. 1994. Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie. Muséum National d'Histoire Naturelle, Paris, 164 p.

Manner, H. I./Mallon, E. 1989. An Annotated list of the vascular plants of Pulawat Atoll. Micronesica 22:23-63.

McCormack, Gerald. 2007. Cook Islands biodiversity and natural heritage. On-line database.

Merlin, M./Juvik, J. 1996. Irá me neeniier nón Chuuk: Plants and their Environments in Chuuk. Program on Environment, East-West Center, Honolulu, Hawai‘i, 121 pp. .

Mune, T. L./Parham, J. W. 1967. The declared noxious weeds of Fiji and their control. Third edition. Department of Agriculture, Fiji. Bulletin No. 48. 87 pp.

Ohba, T. 1994. Flora and vegetation of the northern Mariana Islands, Micronesia. Biological Expedition to the Northern Mariana Islands, Micronesia. In A. Asakua & T. Furuki, eds.  Natural History Research, Special issue no. 1, Chiba, Japan: Natural History Museum and Institute. pp. 13-69.

Orchard, Anthony E., ed. 1993. Flora of Australia. Vol. 50, Oceanic islands 2. Australian Government Publishing Service, Canberra.

Sachet, M.-H. 1983. Botanique de l'ile de Tupai, Iles de la Societe. Atoll Research Bulletin No. 276. Smithsonian Institution, Washington. 26 pp. + photos.

Smith, Albert C. 1981. Flora Vitiensis nova: a new flora of Fiji. National Tropical Botanical Garden, Lawai, Kauai, Hawaii. Volume 2. 810 pp.

Smith, Nicholas M. 2002. Weeds of the wet/dry tropics of Australia - a field guide. Environment Centre NT, Inc. 112 pp.

Space, James C./Imada, Clyde T. 2004. Report to the Republic of Kiribati on invasive plant species on the islands of Tarawa, Abemama, Butaritari and Maiana. Cont. no. 2003-006 to the Pac. Biol. Surv. USDA Forest Service and Bishop Museum, Honolulu. 103 pp.

Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate. 2004. Report to the Government of Niue and the United Nations Development Programme: Invasive plant species on Niue following Cyclone Heta. UNDP NIU/98/G31 - Niue Enabling Activity. 80 pp.

Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar. 2003. Report to the Republic of Palau on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 174 pp.

Stoddart, D. R./Taylor, J. D., eds. 1971. Geography and ecology of Diego Garcia Atoll, Chagos Archipelago. Atoll Research Bulletin No. 149. Smithsonian Institution, Washington. 237 pp.

Stone, Benjamin C. 1970. The flora of Guam. Micronesica 6:1-659.

Swarbrick, John T. 1997. Weeds of the Pacific Islands. Technical paper no. 209. South Pacific Commission, Noumea, New Caledonia. 124 pp.

Sykes, W. R. 1970. Contributions to the flora of Niue. New Zealand Department of Scientific and Industrial Research Bulletin 200, Wellington. 321 pp.

Tassin, Jacques. 2005. Jacques Tassin (IAC-CIRAD), personal communication.

Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. 1994. The flora of Nauru. Smithsonian Institution, Washington. Atoll Research Bulletin 392:1-223.

Thaman, R. R./Tuiwawa, M. 1999. Invasive, potentially invasive and adventive alien plant species of Fiji: a preliminary analysis of their status and measures required for their control. Preliminary draft discussion paper prepared for the SPREP Regional Invasive Species Strategy for the South Pacific Islands Region: Regional Workshop, Nadi, Fiji, 26 September-1 October 1999.

U.S. Dept. Agr., Agr. Res. Serv. 2007. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.

Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawai‘i Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).

Waterhouse, D. F. 1993. The major invertebrate pests and weeds of agriculture in Southeast Asia. The Australian Centre for International Agricultural Research, Canberra. 141 pp.

Waterhouse, D. F. 1997. The major invertebrate pests and weeds of agriculture and plantation forestry in the Southern and Western Pacific. The Australian Centre for International Agricultural Research, Canberra. 93 pp.

Waterhouse, D. F./Norris, K. R. 1987. Biological control: Pacific prospects. Inkata Press, Melbourne. 454 pp.

Welsh, S. L. 1998. Flora Societensis: A summary revision of the flowering plants of the Society Islands. E.P.S. Inc., Orem, Utah. 420 pp.

Wester, Lyndon. 1992. Origin and distribution of adventive flowering plants in Hawai‘i. In: Stone, Charles P.; Smith, Clifford W. and Tunison, J. Timothy. Alien plant invasions in native ecosystems of Hawaii: Management and Research. University of Hawaii, Cooperative National Park Research Studies Unit, Honolulu. University of Hawaii Press. .

Whistler, W. A. 1988. Checklist of the weed flora of western Polynesia. Technical Paper No. 194, South Pacific Commission, Noumea, New Caledonia. 69 pp.

Whistler, W. A. 1990. Ethnobotany of the Cook Islands: The plants, their Maori names, and their uses. Allertonia 5(4):347-424.

Whistler, W. A. 1992. Flora of Tonga [unpublished checklist]. 107 pp.

Wiggins, I. L./Porter, D. M. 1971. Flora of the Galapágos Islands. Stanford University Press. 998 pp.

Yuncker, T. G. 1943. The flora of Niue Island. Bishop Mus. Bull. 178. Bishop Museum Press, Honolulu. 126 pp.

Yuncker, T. G. 1959. Plants of Tonga. Bishop Museum Bull. 220. Bishop Museum Press, Honolulu. 343 pp.