Senna tora (L.) Roxb., Fabaceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Risk assessment results:  Reject, score: 10 (Go to the risk assessment)

Other Latin names:  Cassia tora L.

Common name(s): [more details]

Chamorro: amot-tumaga carabao, mumutong palaéan, mumutun admelon, mumutun palaoan

English: Chinese senna, foetid cassia, Java bean, low senna, peanut weed, sickle senna, sicklepod, stinking cassia

Fijian: kaumoce, kaumothe, pini

French: cassepuante, cassier sauvage, fausse pistache, herbe pistache, pois puant, séné

Hindi: tarota

Japanese: ebisugusa

Samoan: vao pinati, vao pinati

Spanish: bicho, bicho macho, brusca cimarrona, chilinchil, comida de murcielago, ororuz

Tongan: pini papalangi, te‘ekosi, te‘epulu, tengafefeka

Habit:  shrub

Description:  "A small erect glabrous shrub; to 90 cm tall; leaves to 7.5 cm long, with 2-4 pairs of obovate, blunt, or minutely mucronate, ciliate-margined leaflets, each 1-4 cm long; a gland on the rachis between the lowest pair (or lower 2 pairs) of leaflets; flowers paired in axils, yellow, about 9 mm broad, on a 6 mm pedicel; 3 of the stamens infertile; pod very slender, linear, somewhat curved, 12-25 cm long, 3-6 mm wide, beaked, membranous, with broad sutures; seeds about 20-30, nearly 3 mm long, brown, flat" (Stone, 1970; p. 311).

See also Senna occidentalis.

Habitat/ecology:  "It occurs abundantly in open pastures and in pastures under coconuts, and is very common on roadsides and wasteland.  It tolerates fairly low soil fertilities and moisture levels, but grows tallest where conditions are better, when it may form solid stands to the exclusion of all other plants"  (Swarbrick, 1997; p. 15).

In Fiji, "cultivated near sea level and also naturalized on cleared land and becoming a weed in villages and thickets up to an elevation of 900 m"  (Smith, 1985; pp. 110-111). In New Guinea, "a weed of cultivation and roadsides, and particularly troublesome in pastures. In lowland areas with a seasonal climate" (Henty & Pritchard, 1975; p. 115). In Tonga, "an occasional waste-area weed" (Yuncker, 1959; p. 136). In New Caledonia, "largement répandue en lieux incultes bien que tous les spécimens vus soient récents; devient très envahissant dans les pâturages améliorés du Nord-Ouest de la Grande-Terre; le bétail ne la mange pas" (MacKee, 1994; p. 74).

Propagation:  Seed

Native range:  Paleotropical, from India and Ceylon eastward into Polynesia, but not indigenous east of Melanesia and perhaps not there" (Smith, 1985; pp. 110-111).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Tutuila Islands	Tutuila Island		Bishop Museum (Honolulu) (1939) (voucher ID: BISH 153587) Taxon name on voucher: Senna tora (L.) Roxb.
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island		Bishop Museum (Honolulu) (1978) (voucher ID: BISH 652575) Taxon name on voucher: Cassia tora L.
Cook Islands Southern Cook Islands	Ma‘uke Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 15)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced	Glassman, S. F. (1952) (p. 75) Voucher cited: Glassman 2774 (US)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive	Lorence, David H./Flynn, Tim (1998) (p. 21) Naturalized
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Bishop Museum (Honolulu) (1949) (voucher ID: BISH 153634) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Fiji Islands		Bishop Museum (Honolulu) (1860) (voucher ID: BISH 593005) Taxon name on voucher: Cassia tora L.
Fiji Fiji Islands	Kandavu Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Voucher cited: DA 2939
Fiji Fiji Islands	Lakemba Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Voucher cited: Garnock-Jones 896
Fiji Fiji Islands	Matuku Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Voucher cited: Bryan 261
Fiji Fiji Islands	Matuku Island		Bishop Museum (Honolulu) (1924) (voucher ID: BISH 153608) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Rotuma Island		Bishop Museum (Honolulu) (1938) (voucher ID: BISH 614903) Taxon name on voucher: Cassia tora L.
Fiji Fiji Islands	Rotuma Island		Bishop Museum (Honolulu) (1991) (voucher ID: BISH 638037) Taxon name on voucher: Cassia tora L.
Fiji Fiji Islands	Taveuni Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Vouchers cited: Gillespie 4461.4, DA 8923
Fiji Fiji Islands	Taveuni Island		Bishop Museum (Honolulu) (1928) (voucher ID: BISH 153609) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Taveuni Island		Bishop Museum (Honolulu) (1928) (voucher ID: BISH 153610) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Vanua Levu Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Vouchers cited: DA 8745, Greenwood 206B, Krauss 1021
Fiji Fiji Islands	Vanua Levu Island		Bishop Museum (Honolulu) (1966) (voucher ID: BISH 153611) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Vanua Mbalavu Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Voucher cited: DA 10232
Fiji Fiji Islands	Vanua Mbalavu Island		Bishop Museum (Honolulu) (1956) (voucher ID: BISH 32827) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Viti Levu Island	introduced invasive	Smith, Albert C. (1985) (pp. 110-111) Vouchers cited: Greenwood 206, DA 9686, Smith 4500, Degener 14981, DA 5958, DA 9779, DA 10732, DA 9525, DA 3372, DA 11279, H.B.R. Parham 9, Meebold 16462
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1955) (voucher ID: BISH 32828) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1932) (voucher ID: BISH 153604) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1941) (voucher ID: BISH 153605) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1947) (voucher ID: BISH 153606) Taxon name on voucher: Senna tora (L.) Roxb.
Fiji Fiji Islands	Viti Levu Island		Bishop Museum (Honolulu) (1919) (voucher ID: BISH 153607) Taxon name on voucher: Senna tora (L.) Roxb.
French Polynesia French Polynesia Islands	French Polynesia Islands		Swarbrick, John T. (1997) (p. 89)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island		Bishop Museum (Honolulu) (1987) (voucher ID: BISH 540429) Taxon name on voucher: Senna tora (L.) Roxb.
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Welsh, S. L. (1998) (p. 163) Vouchers cited: Florence 3701, BRY 26680
French Polynesia Society Islands	Raiatea (Havai) Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491156) Taxon name on voucher: Cassia tora L.
French Polynesia Society Islands	Tahiti Island	introduced invasive	Welsh, S. L. (1998) (p. 163) Voucher cited: BRY 26035
French Polynesia Society Islands	Tahiti Island		Bishop Museum (Honolulu) (1982) (voucher ID: BISH 492812) Taxon name on voucher: Cassia tora L.
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island		Bishop Museum (Honolulu) (1988) (voucher ID: BISH 560418) Taxon name on voucher: Senna tora (L.) Roxb.
French Polynesia Tuamotu Archipelago	Niau Atoll		Bishop Museum (Honolulu) (1990) (voucher ID: BISH 633898) Taxon name on voucher: Senna tora (L.) Roxb.
Guam Guam Island	Guam Island	introduced invasive	Stone, Benjamin C. (1970) (pp. 311-312) Voucher cited: Stone 4867 (GUAM)
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 99)
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (year unknown) (voucher ID: BISH 153664) Taxon name on voucher: Senna tora (L.) Roxb.
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1936) (voucher ID: BISH 153665) Taxon name on voucher: Senna tora (L.) Roxb.
Guam Guam Island	Guam Island		Bishop Museum (Honolulu) (1953) (voucher ID: BISH 652574) Taxon name on voucher: Cassia tora L.
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 381) Largement répandue en lieux incultes; devient très envahissante dans les pâturages améliorés du Nord-Ouest de la Grande-Terre.
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	MacKee, H. S. (1994) (p. 74) Vouchers cited: Blanchon 797, MacKee 35154, MacKee 39180, Saint-Quentin in MacKee 44936, Auriac in MacKee 44986
Niue Niue	Niue Island	introduced invasive	Whistler, W. A./Atherton, J. (1997) (p. 36)
Niue Niue	Niue Island	introduced invasive	Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 12)
Papua New Guinea Bismarck Archipelago	Bismarck Archipelago	introduced	ILDIS Co-ordinating Centre (2009)
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)	introduced invasive	Henty, E. E./Pritchard, G. H. (1975) (p. 115)
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)	probably native	Waterhouse, D. F. (1997) (p. 65)
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Philippines Philippine Islands	Philippine Islands	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Samoa Western Samoa Islands	Savai‘i Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 71)
Samoa Western Samoa Islands	Savai‘i Island		Bishop Museum (Honolulu) (1931) (voucher ID: BISH 153591) Taxon name on voucher: Senna tora (L.) Roxb.
Samoa Western Samoa Islands	Upolu Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 68)
Samoa Western Samoa Islands	Upolu Island		Bishop Museum (Honolulu) (1968) (voucher ID: BISH 28603) Taxon name on voucher: Senna tora (L.) Roxb.
Samoa Western Samoa Islands	Upolu Island		Bishop Museum (Honolulu) (1929) (voucher ID: BISH 153589) Taxon name on voucher: Senna tora (L.) Roxb.
Samoa Western Samoa Islands	Upolu Island		Bishop Museum (Honolulu) (1921) (voucher ID: BISH 153590) Taxon name on voucher: Senna tora (L.) Roxb.
Solomon Islands Solomon Islands	Solomon Islands	probably native	Swarbrick, John T. (1997) (p. 89)
Solomon Islands Solomon Islands	Solomon Islands	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Solomon Islands Solomon Islands	Solomon Islands	introduced	ILDIS Co-ordinating Centre (2009)
Solomon Islands Solomon Islands	Solomon Islands	native	Hancock, I. R./Henderson, C. P. (1988) (p. 56)
Tonga Ha‘apai Group	Ha‘ano Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 75)
Tonga Ha‘apai Group	Lifuka and Foa Islands	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 75)
Tonga Ha‘apai Group	‘Uiha Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 75)
Tonga Tongatapu Group	‘Eua Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 62)
Tonga Tongatapu Group	Tongatapu Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 56)
Tonga Tongatapu Group	Tongatapu Island		Whistler, W. A. (1992) (p. 32) Vouchers cited: Setchell & Parks 15351, Lawrence 59, Lawrence 116, Soakai 403, Soakai 800
Tonga Tongatapu Group	Tongatapu Island	introduced invasive	Yuncker, T. G. (1959) (p. 136) Vouchers cited: Hülimann 144, Yuncker 15078
Tonga Tongatapu Group	Tongatapu Island		Bishop Museum (Honolulu) (1958) (voucher ID: BISH 41225) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Tongatapu Group	Tongatapu Island		Bishop Museum (Honolulu) (1958) (voucher ID: BISH 41258) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Tongatapu Group	Tongatapu Island		Bishop Museum (Honolulu) (1959) (voucher ID: BISH 41299) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Tongatapu Group	Tongatapu Island		Bishop Museum (Honolulu) (1953) (voucher ID: BISH 153579) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Tongatapu Group	Tongatapu Island		Bishop Museum (Honolulu) (1926) (voucher ID: BISH 153580) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Tongatapu Group	Tongatapu Island		Bishop Museum (Honolulu) (1959) (voucher ID: BISH 403395) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Vava‘u Group	Vava‘u Island	introduced invasive	Space, James C./Flynn, Tim (2001) (p. 69)
Tonga Vava‘u Group	Vava‘u Island		Whistler, W. A. (1992) (p. 32) Voucher cited: Soakai 883
Tonga Vava‘u Group	Vava‘u Island	introduced invasive	Yuncker, T. G. (1959) (p. 136) Voucher cited: Yuncker 16085
Tonga Vava‘u Group	Vava‘u Island		Bishop Museum (Honolulu) (1953) (voucher ID: BISH 153581) Taxon name on voucher: Senna tora (L.) Roxb.
Tonga Vava‘u Group	Vava‘u Island		Bishop Museum (Honolulu) (1959) (voucher ID: BISH 698061) Taxon name on voucher: Senna tora (L.) Roxb.
Vanuatu New Hebrides Islands	Vanuatu (Republic of)		Swarbrick, John T. (1997) (p. 89)
Vanuatu New Hebrides Islands	Vanuatu (Republic of)	introduced invasive	Meyer, Jean-Yves (2000) (p. 107)
Vanuatu New Hebrides Islands	Vanuatu (Republic of)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 168)
Wallis and Futuna Wallis and Futuna (Horne) Islands	Futuna Island	introduced invasive	Morat, Ph./Veillon, J.-M. (1985) As Cassia tora. Morat & Veillon, 1985, cited in Meyer, 2007
Wallis and Futuna Wallis and Futuna (Horne) Islands	Wallis and Futuna Islands		Whistler, W. A. (1988) (p. 27)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia Australia (continental)	Northern Territory	introduced invasive	Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
Australia Australia (continental)	Northern Territory	introduced invasive	Australian Biological Resources Study (2008)
Australia Australia (continental)	Queensland	introduced invasive	Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
Australia Australia (continental)	Queensland	introduced invasive	Australian Biological Resources Study (2008)
Cambodia Cambodia	Cambodia (Kingdom of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Cambodia Cambodia	Cambodia (Kingdom of)	introduced	ILDIS Co-ordinating Centre (2009)
China China	China (People's Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Colombia Colombia	Colombia (Republic of)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
Ecuador (Mainland) Ecuador	Ecuador (Republic of) (continental)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
El Salvador El Salvador	El Salvador (Republic of)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 171)
Honduras Honduras	Honduras (Republic of)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 168)
Indonesia Indonesia	Indonesia (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Indonesia Indonesia	Indonesia (Republic of)	introduced	ILDIS Co-ordinating Centre (2009)
Japan Japan	Japan (country)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
Japan Ryukyu Islands	Ryukyu Islands	introduced	ILDIS Co-ordinating Centre (2009)
South Korea South Korea	South Korea (Republic of)		Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 70)
Malaysia Malaysia	Malaysia (country of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Malaysia Malaysia	Malaysia (country of)	introduced	ILDIS Co-ordinating Centre (2009)
Mexico Mexico	Mexico (United Mexican States)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
Nicaragua Nicaragua	Nicaragua (Republic of)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 168)
Perú Perú	Perú (Republic of)		Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 162)
Taiwan Taiwan	Taiwan Island	native	ILDIS Co-ordinating Centre (2009)
Thailand Thailand	Thailand (Kingdom of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Vietnam Vietnam	Vietnam (Socialist Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Vietnam Vietnam	Vietnam (Socialist Republic of)	introduced	ILDIS Co-ordinating Centre (2009)

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
La Réunion (France) La Réunion Island	La Réunion Island	introduced invasive	Lavergne, Christophe (2006) "Naturalisé"
Maldives Maldive Islands	Maldive Islands	introduced	ILDIS Co-ordinating Centre (2009)
Maldives Maldive Islands	Maldive Islands		Fosberg, F. R. (1957) (p. 20)
Mauritius Mautitius Islands (Mauritius and Rodrigues)	Mauritius Island	introduced	ILDIS Co-ordinating Centre (2009)
Mauritius Mautitius Islands (Mauritius and Rodrigues)	Rodrigues Island	introduced	ILDIS Co-ordinating Centre (2009)
Seychelles Seychelles Islands	Seychelles Islands	introduced	ILDIS Co-ordinating Centre (2009)

Comments:  Especially invasive on the island of ‘Eua, Tonga (Space & Flynn, 2001).

For Galápagos Islands, see S. obtusifolia.

Control: 

Physical:  "Germination may be prevented by mulches if thick enough and undisturbed, but the large seeds give the seedlings great vigor.  Susceptible to hand weeding, chipping and cultivation.  Unpalatable to stock.  Competition from pasture species will probably give control, but it tolerates shade.  Slashing only prevents seeding temporarily"  (Swarbrick, 1997; p. 22).

Chemical:  "Probably susceptible to: 1) residual herbicides at standard rates; 2) translocated herbicides at standard rates, including wick-wiping with glyphosate.  Susceptible to aciflourfen at standard rate.  Probably susceptible to most herbicides suitable for controlling broad-leaved weeds except 'hormones' formulated with butyric acid" (Swarbrick, 1997; p. 22).

Biological: See Waterhouse and Norris (1987) for identification of natural enemies.

Additional information:
Fact sheet from the Government of Queensland, Australia. (PDF format).

Additional online information about Senna tora is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Senna tora as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Senna tora may be available from the Germplasm Resources Information Network (GRIN).

Other Latin names:  Cassia tora L.

References:

Australian Biological Resources Study. 2008. Flora of Australia Online. Australian Biological Resources Study, Canberra.

Bishop Museum (Honolulu). 0. Voucher specimen #BISH 153664 (collector unknown 63).

Bishop Museum (Honolulu). 1860. Voucher specimen #BISH 593005 (Seemann, B.C. 135).

Bishop Museum (Honolulu). 1919. Voucher specimen #BISH 153607 (Greenwood, W. 206).

Bishop Museum (Honolulu). 1921. Voucher specimen #BISH 153590 (Eames, A.J. 86).

Bishop Museum (Honolulu). 1924. Voucher specimen #BISH 153608 (Bryan Jr, E.H. 261).

Bishop Museum (Honolulu). 1926. Voucher specimen #BISH 153580 (Setchell, W.A. 15351).

Bishop Museum (Honolulu). 1928. Voucher specimen #BISH 153609 (Gillespie, J.W. 4461.4).

Bishop Museum (Honolulu). 1928. Voucher specimen #BISH 153610 (Gillespie, J.W. 4461.4).

Bishop Museum (Honolulu). 1929. Voucher specimen #BISH 153589 (Christophersen, E. 434).

Bishop Museum (Honolulu). 1931. Voucher specimen #BISH 153591 (Christophersen, E. 3346).

Bishop Museum (Honolulu). 1932. Voucher specimen #BISH 153604 (Meebold, A. 16462).

Bishop Museum (Honolulu). 1936. Voucher specimen #BISH 153665 (Swezey, O.H.; collector's voucher number unknown).

Bishop Museum (Honolulu). 1938. Voucher specimen #BISH 614903 (St.John, H. 19079).

Bishop Museum (Honolulu). 1939. Voucher specimen #BISH 153587 (Yuncker, T.G. 9361).

Bishop Museum (Honolulu). 1941. Voucher specimen #BISH 153605 (Degener, O. 14981).

Bishop Museum (Honolulu). 1947. Voucher specimen #BISH 153606 (Smith, A.C. 4500).

Bishop Museum (Honolulu). 1949. Voucher specimen #BISH 153634 (Glassman, S.F. 2774).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 153579 (Yuncker, T.G. 15078).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 153581 (Yuncker, T.G. 16085).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 652574 (Fosberg, F.R. 35231).

Bishop Museum (Honolulu). 1955. Voucher specimen #BISH 32828 (Koroiveibau, D. 9686).

Bishop Museum (Honolulu). 1956. Voucher specimen #BISH 32827 (Parham, J.W. 10232).

Bishop Museum (Honolulu). 1958. Voucher specimen #BISH 41225 (Lawrence, E. 59).

Bishop Museum (Honolulu). 1958. Voucher specimen #BISH 41258 (Lawrence, E. 116).

Bishop Museum (Honolulu). 1959. Voucher specimen #BISH 41299 (Soakai, E. 403).

Bishop Museum (Honolulu). 1959. Voucher specimen #BISH 403395 (Soakai, E. 800).

Bishop Museum (Honolulu). 1959. Voucher specimen #BISH 698061 (Soakai, E. 883).

Bishop Museum (Honolulu). 1966. Voucher specimen #BISH 153611 (collector unknown 1021).

Bishop Museum (Honolulu). 1968. Voucher specimen #BISH 28603 (Bristol, M.L. 2071).

Bishop Museum (Honolulu). 1978. Voucher specimen #BISH 652575 (Fosberg, F.R. 58212).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491156 (Florence, J. 3701).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 492812 (Florence, J. 2503).

Bishop Museum (Honolulu). 1987. Voucher specimen #BISH 540429 (Florence, J. 8476).

Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 560418 (Florence, J. 9049).

Bishop Museum (Honolulu). 1990. Voucher specimen #BISH 633898 (Florence 10097).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 638037 (McClatchey, W.C. 214).

Falanruw, Marjorie C./Maka, Jean E./Cole, Thomas G./Whitesell, Craig D. 1990. Common and scientific names of trees and shrubs of Mariana, Caroline, and Marshall Islands. Pac. SW Forest and Range Expt. Stn. Resource Bulletin PSW-67. 91 pp.

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