Pilea microphylla (L.) Liebm., Urticaceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Risk assessment results:  Evaluate, score: 6 (Go to the risk assessment)

Common name(s): [more details]

Chinese: xiao ye leng shui hua

English: artillery fern, artillery plant, gunpowder plant, pistol plant, rockweed

Maori (Cook Islands): kiona topa

Marshallese: likotot tot, loktõktõk, loktôktôk, mõlowi

Niuean: pikimaka

Pingelapan: re

Pohnpeian: limw in tuhke, limwin tuhke

Samoan: tamole, tamole

Satawalese: seling

Tongan: limu

Habit:  herb

Description:  "Annual or short-lived perennial herbs; stems succulent, sometimes slightly woody at base, spreading or cespitose, 5-50 cm long.  Leaves crowded throughout stem length, obovate, elliptic, oblanceolate, or oblong, 2-10 mm long, 1-4 mm wide, those of a pair very unequal in size, sometimes some branches with leaves of a pair equal or all leaves of plant equal, 1-nerved, upper surface with crowded elliptic cystoliths, lower surface finely reticulate, margins entire, petioles 1-4 mm long, always shorter than the blade, stipules ca. 1 mm long.  Flowers unisexual (and the plants monoecious or dioecious), in sessile glomerules or cymes, peduncles 1-6 mm long; staminate flowers 0.6-1 mm long, calyx lobes glabrous, each with a blunt, dorsal, apiculate appendage; pistillate flowers 0.8-1 mm long, outer calyx lobe slightly longer than others, with inconspicuous dorsal, apiculate appendages.  Achenes ellipsoid, compressed, 0.5-1 mm long, slightly longer than the largest calyx lobe" (Wagner et al., 1999; p. 1306).

Habitat/ecology:  "Occasional around houses and on rock walls" (Whistler, 1988; p. 39).

In Hawai‘i, "cultivated and naturalized in low elevation, mesic, disturbed sites" (Wagner et al., 1999; p. 1306).  In Fiji, "a sprawling or prostrate weed, often subligneous at base, on damp rocks and along shaded creeks, often near dwellings" (Smith, 1981; pp. 237-238). In Niue, " a characteristic little succulent plant of rock crevices and fissures in the rather open Lower Terrace forest, where it is often very common.  It also grows in the villages on coral walls and similar places, especially in shade or partial shade.  These habitats are typical for this adventive species in many other tropical Pacific islands"  (Sykes, 1970; pp. 210-211).  In Papua New Guinea, "a weed of nursery and garden beds, paths and house-yards, spreading rapidly on cleared soil in shaded situations" (Henty & Pritchard, 1975; p. 157). In Tonga, "occasional on moist soil, wall, rocks, and so forth, especially about villages" (Yuncker, 1959; p. 102). In New Caledonia, "une forme relativement robuste, est parfois cultivée dans les jardins; une très petite forme est abondante en stations fraîches et humides par terre, sur vieux murs et dans les anfractuosités des terrasses coralliennes du littoral" (MacKee, 1994; p. 137).

Propagation:  Seed. "Small pieces of stem are able to take root, if lying on the soil" (Henty & Pritchard, 1975; p. 157).

Native range:  Southern Florida (US), the West Indies and Mexico south to tropical South America (Wagner et al., 1999; p. 1306).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Manu‘a Islands	Ofu Island		Swarbrick, John T. (1997) (p. 95)
American Samoa Manu‘a Islands	Ofu Island		Whistler, W. A. (1990) Voucher cited: Whistler 3486
American Samoa Manu‘a Islands	Ta‘u Island		Swarbrick, John T. (1997) (p. 95)
American Samoa Manu‘a Islands	Ta‘u Island		Whistler, W. A. (1990) Voucher cited: Whistler 1320
American Samoa Manu‘a Islands	Ta‘u Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1991) (voucher ID: PTBG 10548) Taxon name on voucher: Pilea microphylla
American Samoa Tutuila Islands	Tutuila Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1992) (voucher ID: PTBG 48575) Taxon name on voucher: Pilea microphylla
Australia (Pacific offshore islands) Norfolk Islands	Norfolk Island	introduced invasive	Orchard, Anthony E., ed. (1994) (p. 11) "Recently recorded as a garden weed on bare soil". Voucher cited: W.R. Sykes NI 659 (CHR)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Aguijan (Agiguan, Aguihan) Island	introduced	Raulerson, L. (2006) (p. 27)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1966) (voucher ID: PTBG 23430) Taxon name on voucher: Pilea microphylla
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1982) (voucher ID: PTBG 23980) Taxon name on voucher: Pilea microphylla
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island	introduced	Raulerson, L. (2006) (p. 27)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Tinian Island	introduced	Raulerson, L. (2006) (p. 27)
Cook Islands Northern Cook Islands	Manihiki Atoll		McCormack, Gerald (2011)
Cook Islands Northern Cook Islands	Nassau Island		McCormack, Gerald (2011)
Cook Islands Southern Cook Islands	‘Atiu Island		McCormack, Gerald (2011)
Cook Islands Southern Cook Islands	Mangaia Island		McCormack, Gerald (2011)
Cook Islands Southern Cook Islands	Ma‘uke Island		McCormack, Gerald (2011)
Cook Islands Southern Cook Islands	Miti‘aro Island		McCormack, Gerald (2011)
Cook Islands Southern Cook Islands	Rarotonga Island		McCormack, Gerald (2011)
Ecuador (Galápagos Islands) Isabela Group	Isabela Island	introduced invasive	Charles Darwin Foundation (2008) Observed in survey, no herbarium record.
Ecuador (Galápagos Islands) San Cristóbal Group	San Cristóbal Island	introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Cruz Group	Santa Cruz Island	introduced invasive	Charles Darwin Foundation (2008)
Federated States of Micronesia Chuuk Islands	Chuuk Islands		Swarbrick, John T. (1997) (p. 95)
Federated States of Micronesia Chuuk Islands	Lukunor (Núkúnooch) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Chuuk Islands	Namonuito (Namonweito, Nómwun Wiité) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Kosrae Island	Kosrae Island	introduced invasive	Lorence, David H./Flynn, Timothy (2010) (p. 25) Voucher cited: Lorence 7854 (PTBG) Naturalized
Federated States of Micronesia Kosrae Island	Kosrae Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1996) (voucher ID: PTBG 21010) Taxon name on voucher: Pilea microphylla (L.) Liebm.
Federated States of Micronesia Kosrae Island	Kosrae Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 27724) Taxon name on voucher: Pilea microphylla Liebm.
Federated States of Micronesia Pohnpei Islands	Mokil Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Pohnpei Islands	Nukuoro Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Pohnpei Islands	Pingelap Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Glassman, S. F. (1952) (p. 82) Vouchers cited: Glassman 2453 (US), Glassman 2525 (US), Takamatsu 806 (BISH)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced invasive	Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (pp. 139-140) Vouchers cited: M.J. Balick 4293 (NY, PTBG), S.F. Glassman 2453 (BISH), S.F. Glassman 2525 (BISH), S.F. Glassman 2619 (BISH), S. Perlman 20367 (PTBG), M. Takamatsu 806 (BISH), C.Y.C. Wong 61 (BISH) Naturalized
Federated States of Micronesia Yap Islands	Eauripik Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Fais Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Fais Island	introduced invasive cultivated	Fosberg, F. R./Evans, M. (1969) (p. 6) Voucher cited: Evans 348 (US)
Federated States of Micronesia Yap Islands	Fais Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1965) (voucher ID: PTBG 23289) Taxon name on voucher: Pilea microphylla
Federated States of Micronesia Yap Islands	Faraulep (Faraulap) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Ifalik (Ifaluk) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Satawal Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Satawal Island	introduced invasive	Fosberg, F. R. (1969) (p. 6) Voucher cited: Fosberg 46900 (US)
Federated States of Micronesia Yap Islands	Ulithi Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Woleai Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Federated States of Micronesia Yap Islands	Yap (Waqab) Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1980) (voucher ID: PTBG 24322) Taxon name on voucher: Pilea microphylla
Fiji Fiji Islands	Viti Levu Island	introduced invasive cultivated	Smith, Albert C. (1981) (pp. 237-238) Vouchers cited: Greenwood 1002, 1002B; Meebold 8205; DA 12264; DA 11593
French Polynesia Gambier Islands	Akamaru Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Gambier Islands	Akamaru Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced cultivated	St. John, Harold (1988) (p. 13) Voucher cited: Fosberg 11036 (BISH)
French Polynesia Gambier Islands	Taravai Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Gambier Islands	Taravai Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Society Islands	Huahine Island	introduced invasive	Welsh, S. L. (1998) (p. 290) Voucher cited: Fosberg 63084
French Polynesia Society Islands	Huahine Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Society Islands	Huahine Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence & R. Tahuaitu 11655 (PAP) Adventice
French Polynesia Society Islands	Moorea Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Welsh, S. L. (1998) (p. 290) Voucher cited: Moore 423
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence 3493 (PAP) Adventice
French Polynesia Society Islands	Taha‘a Island	introduced	Fosberg, F. R. (1997) (p. 149)
French Polynesia Society Islands	Tahiti Island	introduced invasive	Welsh, S. L. (1998) (p. 290) Vouchers cited: Grant 4518, Florence 3213, BRY 25968
French Polynesia Society Islands	Tahiti Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Society Islands	Tahiti Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Vouchers cited: J. Florence 3213 (PAP), J. Florence 8757 (PAP) Adventice
French Polynesia Society Islands	Tetiaroa Atoll	introduced cultivated	Sachet, M.-H./Fosberg, F. R. (1983) (p. 65)
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence 3077 (PAP) Adventice
French Polynesia Tuamotu Archipelago	Moruroa Atoll	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Tuamotu Archipelago	Moruroa Atoll	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Tuamotu Archipelago	Raro‘ia Atoll	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Tuamotu Archipelago	Raro‘ia Atoll		Doty, Maxwell S. (1954) (p. 32)
French Polynesia Tuamotu Archipelago	Raro‘ia Atoll	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Tuamotu Archipelago	Takapoto Atoll	introduced cultivated	Sachet, M-H. (1983) (p. 19) Cultivated in pot.
French Polynesia Austral (Tubuai) Islands	Raivavae (Raevavae) Island	introduced invasive cultivated	Florence, J. (1997) (pp. 235-236)
French Polynesia Austral (Tubuai) Islands	Raivavae (Raevavae) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence & W.R. Sykes 11208 (PAP), J. Florence & W.R. Sykes 11326 (PAP) Adventice
French Polynesia Austral (Tubuai) Islands	Rapa Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Austral (Tubuai) Islands	Rimatara (Rimitara) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Austral (Tubuai) Islands	Rurutu Island	introduced invasive cultivated	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Vouchers cited: J. Florence 6221 (PAP), J. Florence 9245 (PAP) Adventice
French Polynesia Austral (Tubuai) Islands	Tubuai Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
Guam Guam Island	Guam Island	introduced invasive	Stone, Benjamin C. (1970) (p. 257) Vouchers cited: Stone 4695 (GUAM), Necker 372, Moore 221, Steeve 137
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Guam Guam Island	Guam Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1987) (voucher ID: PTBG 22834) Taxon name on voucher: Pilea microphylla
Guam Guam Island	Guam Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1966) (voucher ID: PTBG 23879) Taxon name on voucher: Pilea microphylla
State of Hawaii Hawaiian Islands	Hawai‘i (Big) Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1306)
State of Hawaii Hawaiian Islands	Kaua‘i Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1306)
State of Hawaii Hawaiian Islands	Lāna‘i Island	introduced invasive	Oppenheimer, Hank L./Bartlett, Randal T. (2002) (p. 12) Voucher cited: Oppenheimer H109930 (BISH)
State of Hawaii Hawaiian Islands	Maui Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1306)
State of Hawaii Hawaiian Islands	Moloka‘i Island	introduced invasive	Oppenheimer, Hank L. (2006) (p. 14) Voucher cited: Oppenheimer H40407 (BISH, PTBG)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1306)
Kiribati Tungaru (Gilbert) Islands	Butaritari Atoll	introduced invasive	Space, James C./Imada, Clyde T. (2004) (p. 13)
Kiribati Tungaru (Gilbert) Islands	Butaritari Atoll		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1972) (voucher ID: PTBG 33039) Taxon name on voucher: Pilea microphylla
Kiribati Tungaru (Gilbert) Islands	Tarawa Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Kiribati Tungaru (Gilbert) Islands	Tarawa Atoll	introduced invasive	Space, James C./Imada, Clyde T. (2004) (p. 13)
Marshall Islands Ralik Chain	Ailinglaplap (Ailinglapalap) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Marshall Islands Ralik Chain	Ebon (Epoon) Atoll	introduced invasive	Whistler, W. A./Steele, O. (1999) (p. 70)
Marshall Islands Ralik Chain	Jaluit (Jãlwõj) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Marshall Islands Ralik Chain	Jaluit (Jãlwõj) Atoll	introduced invasive	Fosberg, F. R./Sachet, M.-H. (1962) (p. 15) Vouchers cited: St. John 21670 (BISH), Fosberg 26718 (US)
Marshall Islands Ralik Chain	Kili (Kõle) Atoll	introduced invasive	Vander Velde, Nancy/Vander Velde, Brian (2006) (p. 12) "Locally common; ruderal in wetter areas, often growing along with ferns on old stumps ".
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll	introduced invasive	Whistler, W. A./Steele, O. (1999) (p. 70) Vouchers cited: Herbst 8961, Whistler 11260
Marshall Islands Ralik Chain	Kwajalein (Kuwajleen) Atoll		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1998) (voucher ID: PTBG 47252) Taxon name on voucher: Pilea microphylla
Marshall Islands Ralik Chain	Ujelang Atoll	introduced invasive	Whistler, W. A./Steele, O. (1999) (p. 70)
Marshall Islands Ratak Chain	Arno Atoll	introduced invasive	Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 12) "Occasional; on rocks and coconut bases in moist areas".
Marshall Islands Ratak Chain	Majuro (Mãjro) Atoll	introduced invasive	Vander Velde, Nancy (2003) (p. 130)
Marshall Islands Ratak Chain	Mili (Mile) Atoll	introduced cultivated	Vander Velde, Nancy/Vander Velde, Brian (2010) (p. A23) Uncommon; few plants near in private yard, Mili.
Nauru Nauru Island	Nauru Island	introduced invasive	Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 203) Vouchers cited: Fosberg 58713 (US), Thaman & Manner 8 (SUVA)
New Caledonia Îles Loyauté (Loyalte Islands)	Îles Loyauté (Loyalty Islands)	introduced invasive	Tassin, Jacques (2005)
New Caledonia Îles Loyauté (Loyalte Islands)	Île Lifou	introduced invasive cultivated	MacKee, H. S. (1994) (pp. 137-138) Voucher cited: MacKee 5196
New Caledonia Îles Loyauté (Loyalte Islands)	Île Maré	introduced invasive cultivated	MacKee, H. S. (1994) (pp. 137-138) Voucher cited: Schmid 2665
New Caledonia New Caledonia	New Caledonia Islands		Swarbrick, John T. (1997) (p. 95)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	Tassin, Jacques (2005)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive cultivated	MacKee, H. S. (1994) (pp. 137-138) Vouchers cited: Baumann 8447, MacKee 6456, MacKee 9840, Nothis 48 (NOU), MacKee 26222, MacKee 29822
Niue Niue	Niue Island	introduced invasive	Sykes, W. R. (1970) (p. 210) Voucher cited: CHR 169837
Niue Niue	Niue Island	introduced invasive	Whistler, W. A. (1988) (p. 39)
Niue Niue	Niue Island	introduced invasive	Whistler, W. A./Atherton, J. (1997) (p. 48)
Niue Niue	Niue Island	introduced invasive	Yuncker, T. G. (1943) (p. 49) "Common on moist walls and shaded soil".
Niue Niue	Niue Island	introduced invasive	Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 45)
Palau Palau (main island group)	Angaur Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Palau Palau (main island group)	Angaur Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 104)
Palau Palau (main island group)	Angaur Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 60)
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 102)
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 60)
Palau Palau (main island group)	Kayangel (Ngcheangel) Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 8)
Palau Palau (main island group)	Koror Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Palau Palau (main island group)	Koror Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau Palau (main island group)	Koror Island	introduced	Lorence, David H./Flynn, Tim (2010) (p. 44) Voucher cited: Lorence 8263 (PTBG)
Palau Palau (main island group)	Koror Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 60)
Palau Palau (main island group)	Koror Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1998) (voucher ID: PTBG 27941) Taxon name on voucher: Pilea microphylla (L.) Liebm.
Palau Palau (main island group)	Malakal Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau Palau (main island group)	Ngercheu Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108) Carp Resort
Palau Palau (main island group)	Ngercheu Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 60) Carp Resort
Palau Palau (main island group)	Ngerkebesang Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau Palau (main island group)	Peleliu Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 106)
Palau Palau (main island group)	Urukthapel Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Palau Palau (main island group)	Urukthapel Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108)
Palau Rock Islands	Ngerchong Island	introduced invasive	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108)
Palau Rock Islands	Ngerchong Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 60)
Palau Rock Islands	Ngerchong Island		Beleu National Museum (Republic of Palau. Koror.) (1978) (voucher ID: BNM 1433) Taxon name on voucher: Pilea microphylla (L.) Liebm.
Palau Sonsorol Group	Sonsorol Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 61)
Palau Sonsorol Group	Sonsorol Island		Beleu National Museum (Republic of Palau. Koror.) (1970) (voucher ID: BNM 813) Taxon name on voucher: Pilea microphylla (L.) Liebm.
Papua New Guinea Bismarck Archipelago	Bismarck Archipelago	introduced	Peekel, P. G. [translated by E. E. Henty] (1984) (p. 153)
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)	introduced invasive cultivated	Henty, E. E./Pritchard, G. H. (1975) (p. 157) From sea level to 700 m.
Philippines Philippine Islands	Philippine Islands	introduced invasive	Merrill, Elmer D. (1923) (pp. 76-77) In and about towns, on damp walls, etc.
Pitcairn Islands Pitcairn Islands	Pitcairn Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
Samoa Western Samoa Islands	Western Samoa Islands		Whistler, W. A. (1988) (p. 39)
Solomon Islands Solomon Islands	Solomon Islands		Swarbrick, John T. (1997) (p. 95)
Solomon Islands Solomon Islands	Solomon Islands	introduced invasive	Hancock, I. R./Henderson, C. P. (1988) (p. 99)
Tonga Ha‘apai Group	Ha‘apai Islands		Whistler, W. A. (1992) (p. 79) Vouchers cited: Soakai 550, Soakai 695
Tonga Tongatapu Group	Tongatapu Island		Whistler, W. A. (1992) (p. 79) Voucher cited: Soakai 405
Tonga Tongatapu Group	Tongatapu Island	introduced invasive	Yuncker, T. G. (1959) (p. 102) Voucher cited: Yuncker 15174
Tonga Vava‘u Group	Vava‘u Island		Whistler, W. A. (1992) (p. 79) Vouchers cited: Soakai 738, Soakai 1015
Tuvalu Tuvalu (Ellis) Islands	Nui Atoll		Woodroffe, C. D. (1985) (p. 11) Voucher cited: Woodroffe 186 (CHR)
Tuvalu Tuvalu (Ellis) Islands	Tuvalu Islands		Swarbrick, John T. (1997) (p. 95)
United States (other Pacific offshore islands) Johnston Atoll	Johnston Island	introduced invasive	Amerson, A. Binion, Jr./Shelton, Philip C. (1976) (p. 54)
United States (other Pacific offshore islands) Midway Atoll	Eastern Island	introduced invasive	Starr, Forest/Starr, Kim/Loope, Lloyd (2008) (pp. B-11)
United States (other Pacific offshore islands) Midway Atoll	Midway Atoll	introduced invasive	Wagner, Warren L./Herbst, Derral R. (1995) (p. 26) Voucher cited: Herbst & Takeuchi 9069 (BISH)
United States (other Pacific offshore islands) Midway Atoll	Midway Atoll	introduced invasive	Shannon, Robynn K./Wagner, Warren L. (1996) (p. 14) Vouchers cited: Long 1748 (US), Lamoureux 2765 (US)
United States (other Pacific offshore islands) Midway Atoll	Sand Island	introduced invasive	Hughes, Guy D'Oyly (1995) (p. 26) Voucher cited: Herbst & Takeuchi 9069 (BISH)
United States (other Pacific offshore islands) Midway Atoll	Sand Island	introduced invasive	Starr, Forest/Starr, Kim/Loope, Lloyd (2008) (pp. B-11)
United States (other Pacific offshore islands) Palmyra Atoll	Palmyra Atoll	introduced invasive	Wester, Lyndon (1985) (p. 21)
Vanuatu New Hebrides Islands	Vanuatu (Republic of)		Swarbrick, John T. (1997) (p. 95)
Wallis and Futuna Wallis and Futuna (Horne) Islands	Wallis (‘Uvea) Island	introduced invasive	Meyer, Jean-Yves (2007) (p. 38)
Wallis and Futuna Wallis and Futuna (Horne) Islands	Wallis and Futuna Islands		Whistler, W. A. (1988) (p. 39)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
China China	China (People's Republic of)	introduced invasive	Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
China China	Hong Kong	introduced invasive cultivated	Wu, Te-lin (2001) (p. 67) Naturalized, common in Hong Kong.
Colombia Colombia	Colombia (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Costa Rica Costa Rica	Costa Rica (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Ecuador (Mainland) Ecuador	Ecuador (Republic of) (continental)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
El Salvador El Salvador	El Salvador (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Guatemala Guatemala	Guatemala (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Honduras Honduras	Honduras (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Japan Japan	Japan (country)	introduced	Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 181)
Mexico Mexico	Mexico (United Mexican States)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Nicaragua Nicaragua	Nicaragua (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Panama Panama	Panama (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Perú Perú	Perú (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Singapore Singapore	Singapore (Republic of)	introduced invasive	Pallawatta, Nirmalie/Reaser, Jamie K./Gutierrez, Alixis T./eds. (2003) (p. 89)
Singapore Singapore	Singapore (Republic of)	introduced invasive	Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 69) Naturalised

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
British Indian Ocean Territory Chagos Archipelago	Diego Garcia Island		Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 148) Voucher cited: Stoddart 893 (K, US)
British Indian Ocean Territory Chagos Archipelago	Diego Garcia Island	introduced invasive	Whistler, W. Arthur (1996) (p. 30)
British Indian Ocean Territory Chagos Archipelago	Diego Garcia Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 38087) Taxon name on voucher: Pilea microphylla (L.) Liebm.
Maldives Maldive Islands	Malè Atoll		Fosberg, F. R. (1957) (p. 16) "In crevices in walls along streets". Voucher cited: Fosberg 36914
Seychelles Seychelles Islands	Alphonse Island		Robertson, I. A. D./Robertson, S. A./Fosberg, F. R. (1983) (p. 184) Voucher cited: Robertson 2861 (K)
Seychelles Seychelles Islands	Coetivy Island		Robertson, S. A./Fosberg, F. R. (1983) (p. 156) Voucher cited: Robertson 3121 (K)
Seychelles Seychelles Islands	Poivre Island		Robertson, S. A./Fosberg, F. R. (1983) (p. 174) Voucher cited: Robertson 2343 (K)

Additional information:
Additional online information about Pilea microphylla is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Pilea microphylla as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Pilea microphylla may be available from the Germplasm Resources Information Network (GRIN).

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