Pacific Island Ecosystems at Risk (PIER)

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Phyllanthus amarus
Schumach. & Thonn., Phyllanthaceae
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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Other Latin names:  Phyllanthus niruri sensu various Pacific botanists

Common name(s): [more details]

Chamorro: maigo-lalo

Chinese: ku wei ye xia zhu

Chuukese: nikémméwúr, walimo, walumo

English: shatterstone, six o'clock, sleeping plant

I-Kiribati: te kaimatu

Japanese: kidati-kominkaso

Mangarevan: teva

Maori (Cook Islands): moemoe, pī

Marquesan: tuui

Marshallese: jil jino auo, jil jino auö, jil jino awa, jiljino

Palauan: ukalla ruchel

Pohnpeian: limair-poh, limemeirpwong

Samoan: fau lili‘i, fua lili‘i, lau lili‘i, lau lili‘i

Satawalese: walpachi

Tahitian: moemoe, moemoe, moemoe, moemoe uouo, tebe

Tuamotuan: moemoe, moemoe, moemoe, moemoe

Tuvaluan: lakau o iere, te uteute

Ulithian: piapi

Yapese: rurudai

Habit:  herb

Description:  "Small herbs, usually under 30 cm tall, with numerous small oblong-elliptic or squarish leaves, glabrous, about 6-12 mm long; flowers very small, in cymules hidden under the leaves; cymules bisexual, of 1 male and 1 female flower; calyx-lobes 5, acute; pedicels 2 mm long; capsule small, depressed-globose; seeds 5-7-ribbed"  (Stone, 1970; p. 386).

Habitat/ecology:  In Fiji, "occurring at low elevations (not collected above about 50 m) as a weed in villages, gardens, and cultivated fields, often abundantly naturalized" (Smith, 1981). "Common in disturbed places and croplands in Samoa, but uncommon in Tonga"  (Whistler, 1988; p. 19). In New Caledonia, "plante anthropophile, introduite il y a vraisemblablement longtemps, assez commune dans les terrains vagues, à basse altitude, dans la région de Nouméa, d'origine probablement américaine, maintenant pantropicale" (Schmid, 1991; pp. 54, 56-57).

Propagation:  Seed

Native range:  New World.

Presence:

Pacific
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
American Samoa
Manu‘a Islands
Ofu Island   Whistler, W. A. (1990)
Yuncker 9552
American Samoa
Manu‘a Islands
Ofu Island   Yuncker, T. G. (1945) (p. 45)
Voucher cited: Yuncker 9552 (BISH)
American Samoa
Manu‘a Islands
Ofu Island introduced
Ragone, Diane/Lorence, David H. (2003) (p. 52)
American Samoa
Manu‘a Islands
Ofu Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2001) (voucher ID: PTBG 35333)
Taxon name on voucher: Phyllanthus amarus Schumach. & Thonn.
American Samoa
Manu‘a Islands
Ta‘u Island   Whistler, W. A. (1990)
Vouchers cited: Harris 15, Harris 24, Harris 87, Harris 104, Harris 343, Yuncker 9188, Whistler 1312
American Samoa
Manu‘a Islands
Ta‘u Island   Yuncker, T. G. (1945) (p. 45)
Vouchers cited: Yuncker 9188 (BISH), W. Harris 24 (BISH), W. Harris 343 (BISH), W. Harris 15 (BISH), W. Harris 87 (BISH), W. Harris 104 (BISH)
American Samoa
Manu‘a Islands
Ta‘u Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1991) (voucher ID: PTBG 11701)
Taxon name on voucher: Phyllanthus amarus
American Samoa
Swain's Island
Swain's Island introduced
invasive
Whistler, W. A. (1983) (p. 10)
Vouchers cited: Whistler 3402, Bryan 1372, Schultz 12
American Samoa
Tutuila Islands
Tutuila Island   Anon. (1988) (p. 2)
American Samoa
Tutuila Islands
Tutuila Island introduced
invasive
Setchell, William Albert (1924) (pp. 78-79)
As Phyllanthus niruri L.
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Agrigan (Agrihan) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Agrigan (Agrihan) Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Vouchers cited: Fosberg 31433 (US, BISH), Falanruw 2174 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Alamagan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Alamagan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Vouchers cited: Fosberg 31650 (US), Falanruw 1944 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Alamagan Island introduced
Ohba, T. (1994) (p. 59)
Vouchers cited: CBM-BS-58882, CBM-BS-59217
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Anatahan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Anatahan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Voucher cited: Falanruw 1635 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Asuncion Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Asuncion Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Voucher cited: Falanruw 2282 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Farallon de Medinilla (Marpi, Medinilla) Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Maug Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Maug Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Voucher cited: Falanruw 2233? (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Pagan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Pagan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Voucher cited: Anderson 562 (US, BISH)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Rota Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Saipan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Sarigan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Sarigan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 30)
Vouchers cited: Falanruw 1720 (US), Evans 2359 (US, BISH, Fo, NY)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Tinian Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Tinian Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1982) (voucher ID: PTBG 23810)
Taxon name on voucher: Phyllanthus amarus
Cook Islands
Northern Cook Islands
Manihiki Atoll   McCormack, Gerald (2013)
Cook Islands
Northern Cook Islands
Manuae Atoll (Northern Cook Islands)   McCormack, Gerald (2013)
Cook Islands
Northern Cook Islands
Nassau Island   McCormack, Gerald (2013)
Cook Islands
Northern Cook Islands
Penrhyn (Tongareva) Island   McCormack, Gerald (2013)
Cook Islands
Northern Cook Islands
Pukapuka Atoll   McCormack, Gerald (2013)
Cook Islands
Northern Cook Islands
Rakahanga Atoll   McCormack, Gerald (2013)
Cook Islands
Northern Cook Islands
Suwarrow (Taka) Atoll   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Aitutaki Atoll   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
‘Atiu Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Ma‘uke Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Miti‘aro Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Rarotonga Island   McCormack, Gerald (2013)
Federated States of Micronesia
Chuuk Islands
Fefan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Lukunor (Núkúnooch) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Murilo (Mwiriné) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Namoluk (Nómwunuuk) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Namoluk (Nómwunuuk) Atoll   Marshall, Mae/Fosberg, F. R. (1975) (p. 38)
Voucher cited: Marshall 31 (US)
Federated States of Micronesia
Chuuk Islands
Namonuito (Namonweito, Nómwun Wiité) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Pis Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Puluwat (Pwonowót) Atoll introduced
invasive
Manner, H. I./Mallon, E. (1989) (p. 50)
Voucher cited: Manner & Mallon 194 (GUAM)
Federated States of Micronesia
Chuuk Islands
Satawan Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Udot Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Chuuk Islands
Weno (Moen) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Kosrae Island
Kosrae Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Kosrae Island
Kosrae Island introduced
invasive
Lorence, David H./Flynn, Timothy (2010) (p. 22)
Voucher cited: Lorence 7866 (PTBG)
Naturalized
Federated States of Micronesia
Kosrae Island
Kosrae Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1996) (voucher ID: PTBG 21126)
Taxon name on voucher: Phyllanthus amarus Sch. & Th.
Federated States of Micronesia
Kosrae Island
Kosrae Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1982) (voucher ID: PTBG 24270)
Taxon name on voucher: Phyllanthus amarus
Federated States of Micronesia
Pohnpei Islands
Kapingamarangi Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Pohnpei Islands
Mokil (Mwokil) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Pohnpei Islands
Pingelap Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   Glassman, S. F. (1952) (p. 73)
As Phyllanthus niruri L.; vouchers cited: Glassman 2556 (US), Kanehira 735 (NY)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
invasive
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 121)
Vouchers cited: M.J. Balick 4098 (NY, PTBG), M.J. Balick 4303 (NY, PTBG), A. Dores 4 (BISH, NY, PTBG), A. Dores 166 (BISH, NY, PTBG), F.R. Fosberg 58558 (BISH), R. Gallen 112 (NY, PTBG), S.F. Glassman 2556 (BISH), R. Lengsi 20 (NY), D.H. Lorence 9494 (NY, PTBG), P. Raynor 23 (NY, PTBG), S.H. Riesenberg 60 (BISH), H. St. John 21485 (BISH), F. Sohl 15 (NY, PTBG), C.Y.C. Wong 49b (BISH)
Naturalized
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1999) (voucher ID: PTBG 34580)
Taxon name on voucher: Phyllanthus amarus Schumach. & Thonn.
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2001) (voucher ID: PTBG 39143)
Taxon name on voucher: Phyllanthus amarus
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2000) (voucher ID: PTBG 42992)
Taxon name on voucher: Phyllanthus amarus
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2006) (voucher ID: PTBG 44890)
Taxon name on voucher: Phyllanthus amarus
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 46145)
Taxon name on voucher: Phyllanthus amarus
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 48679)
Taxon name on voucher: Phyllanthus amarus
Federated States of Micronesia
Yap Islands
Fais Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Yap Islands
Fais Island introduced
invasive
Fosberg, F. R./Evans, M. (1969) (p. 9)
Vouchers cited: Fosberg 46668 (US), Fosberg 46680 (US), Evans 345 (US)
Federated States of Micronesia
Yap Islands
Faraulep (Faraulap) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Yap Islands
Ifalik (Ifaluk) Atoll   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Federated States of Micronesia
Yap Islands
Lamotrek Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Yap Islands
Satawal Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Yap Islands
Satawal Island introduced
invasive
Fosberg, F. R. (1969) (p. 8)
Voucher cited: Fosberg 46871 (US)
Federated States of Micronesia
Yap Islands
Ulithi Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Yap Islands
Woleai Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Federated States of Micronesia
Yap Islands
Yap (Waqab) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Fiji
Fiji Islands
Lakemba Island introduced
invasive
Smith, Albert C. (1981) (p. 465)
Voucher cited: Garnock-Jones 909
Fiji
Fiji Islands
Vanua Levu Island introduced
invasive
Smith, Albert C. (1981) (p. 465)
Voucher cited: DA 8727
Fiji
Fiji Islands
Vanua Mbalavu Island introduced
invasive
Smith, Albert C. (1981) (p. 465)
Voucher cited: DA 10233
Fiji
Fiji Islands
Viti Levu Island introduced
invasive
Smith, Albert C. (1981) (p. 465)
Vouchers cited: Greenwood 208, DA 10330, DA 8562, DA 9690, DA 11320, Smith 9498, DA 11001, DA 10045, DA 9883, DA 409, DA 1687
French Polynesia
Clipperton Island
Clipperton (Arue, Avera) Island   Sachet, M.-H. (1962) (p. 93)
French Polynesia
Gambier Islands
Agakauitai Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Gambier Islands
Agakauitai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Agakauitai Island introduced
St. John, Harold (1988) (p. 17)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Voucher cited: St. John 14911 (BISH)
French Polynesia
Gambier Islands
Akamaru Island introduced
St. John, Harold (1988) (p. 18)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Observed
French Polynesia
Gambier Islands
Aukena Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Gambier Islands
Aukena Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Aukena Island introduced
St. John, Harold (1988) (p. 17)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Voucher cited: St. John 14624 (BISH)
French Polynesia
Gambier Islands
Makaroa Island introduced
St. John, Harold (1988) (p. 18)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Observed
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
St. John, Harold (1988) (p. 17)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Vouchers cited: Agassiz 110 (GH), Agassiz 111 (GH), Fosberg 11035 (BISH)
French Polynesia
Gambier Islands
Taravai Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Gambier Islands
Taravai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Taravai Island introduced
St. John, Harold (1988) (p. 17)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Voucher cited: St. John 14772 (BISH)
French Polynesia
Gambier Islands
Tauna Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Gambier Islands
Tauna Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Tauna Island introduced
St. John, Harold (1988) (p. 18)
Phyllanthus debilis auct.: H. St. John non J.T. Klein ex C.L. Willdenow. Voucher cited: St. John 14755 (BISH)
French Polynesia
Gambier Islands
Temoe Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Gambier Islands
Temoe Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Totegegie Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Eiao Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Eiao Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: K.R. Wood 4485 (PAP)
Adventice
French Polynesia
Marquesas Islands
Fatu Hiva Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 21712)
Taxon name on voucher: Phyllanthus amarus Schum. & Thonn.
French Polynesia
Marquesas Islands
Hatuta'a (Hatutu) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Hatuta'a (Hatutu) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Hatuta'a (Hatutu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Hiva Oa Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39748)
Taxon name on voucher: Phyllanthus amarus
French Polynesia
Marquesas Islands
Hiva Oa Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 41674)
Taxon name on voucher: Phyllanthus amarus
French Polynesia
Marquesas Islands
Mohotani (Mohotane, Mohotane) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Mohotani (Mohotane, Mohotane) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 4110 (PAP), J. Florence 6699 (PAP), J. Florence 8543 (PAP)
Adventice
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39396)
Taxon name on voucher: Phyllanthus amarus
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 4445 (PAP)
Adventice
French Polynesia
Society Islands
Bora Bora Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Bora Bora Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 12053 (PAP), J. Florence 12150 (PAP)
Adventice
French Polynesia
Society Islands
Huahine Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Maiao (Maiaoiti, Mauiti, Tupuaemanu) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Maiao (Maiaoiti, Mauiti, Tupuaemanu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Maupiti (Maurua) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Maupiti (Maurua) Island introduced
invasive
Fosberg, F. R./Sachet, M.-H. (1987) (p. 43)
Vouchers cited: Raynal 17867, Fosberg 64804 (US), Fosberg 64909 (US), Fosberg & Sachet 64964 (US)
French Polynesia
Society Islands
Maupiti (Maurua) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Moorea Island introduced
invasive
Welsh, S. L. (1998) (p. 112)
Voucher cited: Florence 2203
French Polynesia
Society Islands
Moorea Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Moorea Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 2203 (PAP), J. Florence 8242 (PAP)
Adventice
French Polynesia
Society Islands
Mopelia (Mapetia, Maupiha‘a, Maupihoa, Mopiha‘a) Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Mopelia (Mapetia, Maupiha‘a, Maupihoa, Mopiha‘a) Atoll introduced
invasive
Sachet, M.-H. (1983) (p. 29)
Voucher cited: Sachet 954 (US)
French Polynesia
Society Islands
Mopelia (Mapetia, Maupiha‘a, Maupihoa, Mopiha‘a) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Welsh, S. L. (1998) (p. 112)
Voucher cited: Moore 31
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 3396 (PAP)
Adventice
French Polynesia
Society Islands
Taha‘a Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Taha‘a Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 5269 (PAP)
Adventice
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Welsh, S. L. (1998) (p. 112)
Voucher cited: Grant 3646
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Tetiaroa Atoll introduced
invasive
Welsh, S. L. (1998) (p. 112)
Voucher cited: Florence 3259
French Polynesia
Society Islands
Tetiaroa Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Tetiaroa Atoll introduced
invasive
Sachet, M.-H./Fosberg, F. R. (1983) (p. 45)
Vouchers cited: Sachet `651 (US), Fosberg 63784 (US), Fosberg 54595 (US), Raynal 18023 (P)
French Polynesia
Society Islands
Tetiaroa Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 3259 (PAP)
Adventice
French Polynesia
Society Islands
Tupai Island introduced
invasive
Welsh, S. L. (1998) (p. 112)
Voucher cited: Grant 4828
French Polynesia
Society Islands
Tupai Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Society Islands
Tupai Island introduced
invasive
Sachet, M.-H. (1983) (p. 19)
Vouchers cited: Sachet 1989 (US), Grant 4828 (BISH)
French Polynesia
Society Islands
Tupai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Ana‘a Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Ana‘a Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Hao (Hau) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Makatea (Ma‘atea) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Makatea (Ma‘atea) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 3027 (PAP), J. Florence 3099 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Manihi Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Manihi Atoll introduced
invasive
Sachet, M-H. (1983) (p. 27)
Voucher cited: Brooks & Brooks 20 (BISH)
French Polynesia
Tuamotu Archipelago
Manihi Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Moruroa Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Niau Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Niau Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Niau Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1990) (voucher ID: PTBG 30716)
Taxon name on voucher: Phyllanthus amarus Schumach. & Thonn.
French Polynesia
Tuamotu Archipelago
Rangiroa (Rairoa) Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Rangiroa (Rairoa) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Raro‘ia Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Raro‘ia Atoll   Doty, Maxwell S. (1954) (p. 32)
French Polynesia
Tuamotu Archipelago
Raro‘ia Atoll introduced
invasive
abo. intro.
Meyer, Jean-Yves/Fourdrigniez, Marie (2008) (p. 14)
Adventice d'introduction polynésienne.
French Polynesia
Tuamotu Archipelago
Raro‘ia Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Takapoto Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Takapoto Atoll introduced
invasive
Sachet, M-H. (1983) (p. 27)
Voucher cited: Sachet 2044 (US)
French Polynesia
Tuamotu Archipelago
Takapoto Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Tenarunga Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Tikehau (Tikahau) Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Tikehau (Tikahau) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 7101 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Toau (Tovau) Atoll introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Tuamotu Archipelago
Toau (Tovau) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Vahanga Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Austral (Tubuai) Islands
Raivavae (Raevavae) Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Austral (Tubuai) Islands
Raivavae (Raevavae) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rimatara (Rimitara) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
invasive
Florence, J. (1997) (pp. 120-122)
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 6057 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Tubuai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
Guam
Guam Island
Guam Island introduced
invasive
Stone, Benjamin C. (1970) (p. 386)
Voucher cited: Stone 3984 (GUAM)
Guam
Guam Island
Guam Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Japan (offshore islands)
Ogasawara (Bonin) Islands
Bonin (Ogasawara) Islands introduced
Kato, Hidetoshi (2007)
Kiribati
Line Islands
Caroline Atoll   Clapp, R. B./Sibley, F. C. (1971) (p. 8)
Voucher cited: Long 3283 (HAW)
Kiribati
Line Islands
Flint Island introduced
invasive
St. John, Harold/Fosberg, F. Raymond (1937) (p. 4)
Voucher cited: St. John & Fosberg 17459 (BISH)
Kiribati
Line Islands
Kiritimati (Christmas) Atoll   Chock, Alvin K./Hamilton, Jr., Dean C. (1962) (p. 5)
Vouchers cited: Bergman 16, Fosberg 13276 (BISH)
Kiribati
Line Islands
Kiritimati (Christmas) Atoll introduced
invasive
Wester, Lyndon (1985) (p. 26)
Kiribati
Line Islands
Kiritimati (Christmas) Atoll introduced
invasive
Christophersen, Erling (1927) (pp. 25-26, 78)
Voucher cited: Bergman 16
Kiribati
Line Islands
Tabuaeran (Fanning) Atoll introduced
invasive
Wester, Lyndon (1985) (p. 26)
Kiribati
Line Islands
Tabuaeran (Fanning) Atoll introduced
invasive
Christophersen, Erling (1927) (p. 42)
Lawn and waste places near the settlement and Cable Station. Voucher cited: Ball 2
Kiribati
Line Islands
Teraina (Washington) Island introduced
invasive
Wester, Lyndon (1985) (p. 26)
Kiribati
Line Islands
Teraina (Washington) Island introduced
invasive
Christophersen, Erling (1927) (pp. 56, 78)
In lawns and waste places at the settlement. Voucher cited: Bergman 75
Kiribati
Phoenix Islands
Kanton Island introduced
invasive
Fosberg, F. R./Stoddart, D. R. (1994) (pp. 25-26)
Vouchers cited: Fosberg & Stoddart 54715 (US), Clapp P-71-25 (US), Degener & Degener 24646 (BISH)
Kiribati
Phoenix Islands
Nikumaroro (Gardner) Atoll introduced
invasive
Fosberg, F. R./Stoddart, D. R. (1994) (pp. 25-26)
Woodward & Long 2471a (US)
Kiribati
Phoenix Islands
Orona (Hull) Atoll introduced
invasive
Fosberg, F. R./Stoddart, D. R. (1994) (pp. 25-26)
Vouchers cited: Fosberg & Stoddart 54803 (US, HAW, K), Llong 2005 (US), Clegern 79 (BISH)
Kiribati
Tungaru (Gilbert) Islands
Abemama Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 76)
Voucher specimens collected: Imada 2003-137 (BISH 697749)
Kiribati
Tungaru (Gilbert) Islands
Butaritari Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 77)
Kiribati
Tungaru (Gilbert) Islands
Butaritari Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1972) (voucher ID: PTBG 31524)
Taxon name on voucher: Phyllanthus amarus
Kiribati
Tungaru (Gilbert) Islands
Maiana Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 79)
Voucher: Imada 2003-83 (BISH 697698)
Kiribati
Tungaru (Gilbert) Islands
Nonouti Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Kiribati
Tungaru (Gilbert) Islands
Onotoa Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Kiribati
Tungaru (Gilbert) Islands
Onotoa Atoll   Moul, Edwin T. (1957) (p. 15)
Vouchers cited: Moul 8087 (US), Moul 8148 (US), Moul 8231 (US)
Kiribati
Tungaru (Gilbert) Islands
Tabiteuea Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Kiribati
Tungaru (Gilbert) Islands
Tarawa Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 75)
Voucher: Imada 2003-22 (BISH 697637)
Kiribati
Tungaru (Gilbert) Islands
Tarawa Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1972) (voucher ID: PTBG 31525)
Taxon name on voucher: Phyllanthus amarus
Marshall Islands
Ralik Chain
Ailinglaplap (Ailinglapalap) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Bikini (Pikinni) Atoll introduced
Fosberg, F. R. (1988) (p. 13)
Marshall Islands
Ralik Chain
Ebon (Epoon) Atoll   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll introduced
invasive
St. John, Harold (1960) (pp. 329-330)
Vouchers cited: St. Joh 23706, Lane 57-105
Marshall Islands
Ralik Chain
Jaluit (Jãlwõj) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Jaluit (Jãlwõj) Atoll introduced
invasive
Fosberg, F. R./Sachet, M.-H. (1962) (p. 24)
Vouchers cited: Fosberg 26719 (US), Fosberg 41394 (US)
Marshall Islands
Ralik Chain
Kili (Kõle) Atoll introduced
invasive
Vander Velde, Nancy/Vander Velde, Brian (2006) (p. 8)
"Common; in fields, around houses and ruderal areas".
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
invasive
Whistler, W. A./Steele, O. (1999) (p. 50)
Vouchers cited: Bryan s.n., Fosberg 26487, Fosberg 31173, Fosberg 36643, Herbst 8886, Herbst 8916, Herbst 8945, Herbst 8969, Herbst 9002, Whistler 11262
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Vouchers cited: Fosberg 31173 (US), Fosberg 26487 (US)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1998) (voucher ID: PTBG 47458)
Taxon name on voucher: Phyllanthus amarus
Marshall Islands
Ralik Chain
Lae Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Lae Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Voucher cited: Fosberg 34092 (US)
Marshall Islands
Ralik Chain
Namorik (Namdik) Atoll   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Marshall Islands
Ralik Chain
Ujae Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Ujae Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Voucher cited: Fosberg 34306 (US)
Marshall Islands
Ralik Chain
Ujelang Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ralik Chain
Ujelang Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Voucher cited: Fosberg 34183 (US)
Marshall Islands
Ralik Chain
Wotho (Wõtto) Atoll   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Marshall Islands
Ratak Chain
Arno Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ratak Chain
Arno Atoll   Anderson, Donald (1951) (p. v)
Voucher cited: Anderson 3627
Marshall Islands
Ratak Chain
Arno Atoll introduced
invasive
Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 7)
"Common to locally abundant; weed in farms, roadsides and ruderal areas".
Marshall Islands
Ratak Chain
Jemo Island introduced
Fosberg, F. Raymond (1990) (p. 36)
Marshall Islands
Ratak Chain
Jemo Island introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Voucher cited: Fosberg 33869 (US)
Marshall Islands
Ratak Chain
Likiep Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ratak Chain
Likiep Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Vouchers cited: Fosberg 33832 (US), Fosberg 27015 (US)
Marshall Islands
Ratak Chain
Majuro (Mãjro) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ratak Chain
Majuro (Mãjro) Atoll introduced
invasive
Vander Velde, Nancy (2003) (p. 95)
Marshall Islands
Ratak Chain
Maloelap Atoll   Imada, Clyde (2006)
Marshall Islands
Ratak Chain
Mili (Mile) Atoll introduced
invasive
Vander Velde, Nancy/Vander Velde, Brian (2010) (p. A15)
Common; in fields and ruderal areas, Mili.
Marshall Islands
Ratak Chain
Utirik (Utrik, Utrõk) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Marshall Islands
Ratak Chain
Utirik (Utrik, Utrõk) Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 13)
Voucher cited: Fosberg 33688 (US)
Nauru
Nauru Island
Nauru Island introduced
invasive
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 134)
Vouchers cited: Fosberg 58685 (US), Fosberg 58733 (US), Scully 155N (US?), Thaman & Manner 47 (SUVA), Swarbrick 9571 (BRIU)
New Caledonia
Îles Loyauté (Loyalte Islands)
Île Walpole introduced
MacKee, H. S. (1994) (pp. 53-54)
Voucher cited: Cherrier in MacKee 44084
New Caledonia
New Caledonia
New Caledonia Islands   Swarbrick, John T. (1997) (p. 95)
New Caledonia
New Caledonia Archipelago
Île Grande Terre introduced
invasive
Tassin, Jacques (2005)
New Caledonia
New Caledonia Archipelago
Île Grande Terre introduced
MacKee, H. S. (1994) (pp. 53-54)
Vouchers cited: Däniker 1212 (Z), Baumann 5416, MacKee 7838, MacKee 12162, MacKee 21865, MacKee 33129
Niue
Niue
Niue Island introduced
invasive
Sykes, W. R. (1970) (p. 95)
P. niruri sensu auct. non L. in Sykes. Voucher cited: CHR 169733
Niue
Niue
Niue Island introduced
invasive
Whistler, W. A. (1988) (p. 19)
Niue
Niue
Niue Island introduced
invasive
Whistler, W. A./Atherton, J. (1997) (p. 33)
Niue
Niue
Niue Island introduced
invasive
Yuncker, T. G. (1943) (p. 72)
As Phyllanthus niruri L. Voucher cited: Yuncker 9907 (BISH)
"Common in moist soil of gardens and plantations".
Niue
Niue
Niue Island introduced
invasive
Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 45)
Palau
Hatohobei Group
Helen Reef introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 110)
Palau
Hatohobei Group
Tobi Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 110)
Palau
Kayangel Atoll
Kayangel Islet introduced
invasive
Miles, Joel (2015) (p. 2)
ref cites other taxa as "Cultivated," but not this one, so implication is that this taxon was found in a non-cultivated situation (PT@PhilipT.com/20170602)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 104)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Voucher: Lorence 9734 (PTBG, BISH, BNM, NY, TAMU, US)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 224)
Taxon name on voucher: Phyllanthus amarus
Palau
Palau (Belau ) (main island group)
Babeldaob Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 102)
Palau
Palau (Belau ) (main island group)
Kayangel (Ngcheangel) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Palau
Palau (Belau ) (main island group)
Kayangel (Ngcheangel) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 105)
Palau
Palau (Belau ) (main island group)
Kayangel (Ngcheangel) Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Voucher: Lorence 9782 (PTBG, BNM, NY, US)
Palau
Palau (Belau ) (main island group)
Kayangel (Ngcheangel) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 75)
Taxon name on voucher: Phyllanthus amarus Schumach. & Thonn.
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   Beleu National Museum (Republic of Palau. Koror.) (1976) (voucher ID: BNM 10027)
Taxon name on voucher: Phyllanthus amarus Schum. & Thonn.
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   Beleu National Museum (Republic of Palau. Koror.) (1968) (voucher ID: BNM 351)
Taxon name on voucher: Phyllanthus amarus Sch. & Thonn.
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   Beleu National Museum (Republic of Palau. Koror.) (1976) (voucher ID: BNM 10029)
Taxon name on voucher: Hedyotis corymbosa (L.) Lam.
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1998) (voucher ID: PTBG 27937)
Taxon name on voucher: Phyllanthus amarus Schum. & Thonn.
Palau
Palau (Belau ) (main island group)
Malakal (Ngemelachel) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Palau
Palau (Belau ) (main island group)
Malakal (Ngemelachel) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau
Palau (Belau ) (main island group)
Ngercheu Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108)
Carp Resort
Palau
Palau (Belau ) (main island group)
Ngercheu Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Carp Resort
Palau
Palau (Belau ) (main island group)
Ngerkebesang Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau
Palau (Belau ) (main island group)
Peleliu (Beliliou) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 106)
Palau
Palau (Belau ) (main island group)
Peleliu (Beliliou) Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Voucher: Lorence 9768 (PTBG, BNM, NY, US)
Palau
Palau (Belau ) (main island group)
Peleliu (Beliliou) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 60)
Taxon name on voucher: Phyllanthus amarus Schumach. & Thonn.
Palau
Rock Islands (Chelbacheb)
Bkul a Chesemiich Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Palau
Rock Islands (Chelbacheb)
Bkul a Chesemiich Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108)
Palau
Rock Islands (Chelbacheb)
Ngerchong Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108)
Palau
Rock Islands (Chelbacheb)
Ngerchong Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Palau
Rock Islands (Chelbacheb)
Ngerchong Island   Beleu National Museum (Republic of Palau. Koror.) (1978) (voucher ID: BNM 1220)
Taxon name on voucher: Phyllanthus amarus Sch. & Th.
Palau
Sonsorol Group
Fanna Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 143)
Palau
Sonsorol Group
Fanna Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109)
Palau
Sonsorol Group
Merir Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109)
Palau
Sonsorol Group
Pulo Ana Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109)
Palau
Sonsorol Group
Sonsorol Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109)
Palau
Sonsorol Group
Sonsorol Island   Beleu National Museum (Republic of Palau. Koror.) (1968) (voucher ID: BNM 356)
Taxon name on voucher: Phyllanthus amarus Sch. & Th.
Papua New Guinea
Bismarck Archipelago
Bismarck Archipelago   Peekel, P. G. [translated by E. E. Henty] (1984) (p. 295)
Generally common, on lawns and in gardens.
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island) native
Henty, E. E./Pritchard, G. H. (1975) (p. 102)
Widespread and common at low altitudes.
Philippines
Philippine Islands
Philippine Islands   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 276)
Philippines
Philippine Islands
Philippine Islands   Merrill, Elmer D. (1923) (p. 394)
As Phyllanthus nirui Linn. At low and medium altitudes, a common weed.
Samoa
Western Samoa Islands
Western Samoa Islands   Whistler, W. A. (1988) (p. 19)
Solomon Islands
Solomon Islands
Solomon Islands   Swarbrick, John T. (1997) (p. 95)
Tonga
Ha‘apai Group
Ha‘apai Islands   Whistler, W. A. (1992) (p. 25)
Voucher cited: Soakai 540
Tonga
Ha‘apai Group
Lifuka Island   Yuncker, T. G. (1959) (p. 159)
Voucher cited: Yuncker 15719
Tonga
Tonga Islands
Tonga Islands   Whistler, W. A. (1992) (p. 25)
Voucher cited: Soakai 595
Tonga
Tongatapu Group
Tongatapu Island   Yuncker, T. G. (1959) (p. 159)
Voucher cited: Setchell & Parks 15329
Tonga
Vava‘u Group
Niuatoputapu Island   Yuncker, T. G. (1959) (p. 159)
Voucher cited: Hürlimann 329
Tonga
Vava‘u Group
Vava‘u Island   Whistler, W. A. (1992) (p. 25)
Voucher cited: Crosby 154
Tuvalu
Tuvalu (Ellis) Islands
Nui Atoll   Woodroffe, C. D. (1985) (p. 13)
Voucher cited: Woodroffe 105 (CHR)
Tuvalu
Tuvalu (Ellis) Islands
Tuvalu Islands   Swarbrick, John T. (1997) (p. 95)
United States (other Pacific offshore islands)
Baker Island
Baker Island introduced
invasive
Christophersen, Erling (1927) (pp. 71, 78)
Voucher cited: Christophersen 28
United States (other Pacific offshore islands)
Palmyra Atoll
Palmyra Atoll introduced
invasive
Wester, Lyndon (1985) (p. 26)
United States (other Pacific offshore islands)
Wake Islands
Wake Island introduced
invasive
Fosberg, F. R. (1959) (p. 15)
Reported as a weed by Bryan (1942) as P. niruri, probably P. amarus but not specimen not seen.
United States (other Pacific offshore islands)
Wake Islands
Wake Island introduced
Fosberg, F. R./Sachet, M.-H. (1969) (p. 11)
Voucher cited: Fosberg 43524 (US)
Vanuatu
New Hebrides Islands
Vanuatu (Republic of)   Swarbrick, John T. (1997) (p. 95)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Futuna Island introduced
invasive
Morat, Ph./Veillon, J.-M. (1985)
Morat & Veillon, 1985, cited in Meyer, 2007
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis (‘Uvea) Island introduced
invasive
Meyer, Jean-Yves (2007) (p. 33)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis and Futuna Islands   Whistler, W. A. (1988) (p. 19)
Pacific Rim
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Australia
Australia (continental)
Australia (continental)   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 276)
China
China
China (People's Republic of) introduced
invasive
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2013)
Dry fields, roadsides, wastelands, forest margin, scrubby woods; below 100-600 m. Guangdong, Guangxi, Hainan, Yunnan.
Colombia
Colombia
Colombia (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Costa Rica
Costa Rica
Costa Rica (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Ecuador (Mainland)
Ecuador
Ecuador (Republic of) (continental) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
El Salvador
El Salvador
El Salvador (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Guatemala
Guatemala
Guatemala (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Honduras
Honduras
Honduras (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Indonesia
Indonesia
Indonesia (Republic of)   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 276)
Malaysia
Malaysia
Malaysia (country of)   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 276)
Mexico
Mexico
Mexico (United Mexican States) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Nicaragua
Nicaragua
Nicaragua (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Panama
Panama
Panama (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Perú
Perú
Perú (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Singapore
Singapore
Singapore (Republic of) introduced
invasive
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 69)
Naturalised
Taiwan
Taiwan Island
Taiwan Island introduced
invasive
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2013)
Dry fields, roadsides, wastelands, forest margin, scrubby woods; below 100-600 m.
Thailand
Thailand
Thailand (Kingdom of)   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 276)
Vietnam
Vietnam
Vietnam (Socialist Republic of)   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 276)
Indian Ocean
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Australia (Indian Ocean offshore islands)
Christmas Island Group
Christmas Island introduced
invasive
Swarbrick, J. T. (1997) (p. 123)
Australia (Indian Ocean offshore islands)
Cocos (Keeling) Islands
Cocos (Keeling) Islands introduced
invasive
Orchard, Anthony E., ed. (1993) (p. 37)
British Indian Ocean Territory
Chagos Archipelago
Diego Garcia Island introduced
invasive
Topp, J. M. W. (1988) (p. 13)
British Indian Ocean Territory
Chagos Archipelago
Diego Garcia Island introduced
Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 153)
Vouchers cited: Bourne 42 (K), Stoddart 894 (K), Rhyne 674 (US)
British Indian Ocean Territory
Chagos Archipelago
Diego Garcia Island introduced
invasive
Whistler, W. Arthur (1996) (p. 27)
British Indian Ocean Territory
Chagos Archipelago
Diego Garcia Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 37915)
Taxon name on voucher: Phyllanthus amarus Schumach. & Thonn.
La Réunion (France)
La Réunion Island
La Réunion Island introduced
invasive
MacDonald, I. A. W./Thebaud, C./Strahm, W. A./Strasberg, D. (1991) (pp. 51-61)
Maldives
Maldive Islands
Malè Atoll   Fosberg, F. R. (1957) (p. 24)
"Occasional in weedy sod back of beach". Voucher cited: Fosberg 36861 (US)
Seychelles
Seychelles Islands
Alphonse Island   Robertson, I. A. D./Robertson, S. A./Fosberg, F. R. (1983) (p. 181)
Voucher cited: Robertson 2852 (K)
Seychelles
Seychelles Islands
Coetivy Island   Robertson, S. A./Fosberg, F. R. (1983) (p. 149)
Voucher cited: Robertson 3083 (K)
Seychelles
Seychelles Islands
Cousin Island introduced
invasive
Fosberg, F. R. (1983) (p. 28)
Vouchers cited: Fosberg 52082 (US), Fosberg 52121 (US)
Seychelles
Seychelles Islands
Desnoeufs Island   Wilson, J. R. (1983) (p. 209)
Seychelles
Seychelles Islands
Frégate Island   Robertson, S. A./Todd, D. M. (1983) (p. 51)
Voucher cited: Robertson 2713
Seychelles
Seychelles Islands
Marie-Louise Island   Wilson, J. R. (1983) (p. 191)
Seychelles
Seychelles Islands
Platte Island   Robertson, S. A./Fosberg, F. R. (1983) (p. 161)
Voucher cited: Robertson 3000 (K)
Seychelles
Seychelles Islands
Poivre Island   Robertson, S. A./Fosberg, F. R. (1983) (p. 170)
Voucher cited: Robertson 2345 (K)

Comments:  Similar to P. debilis and P. niruri (Stone, 1970; p. 386).


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This page was created on 26 JUL 2002 and was last updated on 24 JAN 2007.