Persea americana Mill., Lauraceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Risk assessment results:  Low risk (based on second screen), score: 3 (Go to the risk assessment)

Common name(s): [more details]

Chamorro: alageta

English: alligator pear, avocado, avocado pear

French: avocat, avocatier, avogado

Maori (Cook Islands): ‘āpoka, ‘āpuka, ‘āvōta

Marquesan: avoka, avoka

Niuean: avoka

Palauan: bata

Samoan: ‘avoka

Spanish: aguacate, palto

Tahitian: avoka, avoka, avoka, avota

Tongan: ‘avoka

Habit:  tree

Description:  "Trees; young branches terete, glabrate, often with conspicuous leaf scars.  Leaves dark green on upper surface, lower surface pale green, alternate, variable in shape and size, but usually elliptic, 10-25 cm long, 5-10 cm wide, margins entire.  Flowers in pubescent inflorescences shorter than leaves, clustered at the tips of the branches or on short, lateral young branches; tepals equal, ca. 5 mm long, densely pubescent; fertile stamens 9, staminodia present; anthers tetrathecal; ovary pubescent.  Fruit large, pear-shaped, up to 10 cm or more long, not subtended by persistent tepals, cupule absent" (Wagner et al., 1999; pp. 847-848).

Habitat/ecology:  Trees grow from sea level up to 2250 m altitude. It cannot stand waterlogging. Branches are easily damaged by wind. It needs to be in a frost-free location or where frosts are rare. (Ecoport)

In Hawai‘i, "naturalized in disturbed mesic areas, 10-66 m" (Wagner et al., 1999; pp. 847-848). Moist uplands in the Galápagos Islands (McMullen, 1999; p. 64).

Propagation:  Seed. Spread by cattle and pigs in the Galapagos Islands (McMullen, 1999; p. 64).

Native range:  Central America, widely cultivated.

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Chile (Rapa Nui Island) Rapa Nui (Easter) Island	Rapa Nui (Easter) Island (Isla de Pasqua)	introduced invasive	Meyer, Jean-Yves (2008) (p. 27) "Considéré comme « envahissant dans les milieux naturels » par J. Tassin, 2005. En effet, cette espèce est naturalisée (à Ana Tepahu ou dans le Rano Kau par exemple) mais sans former des couverts denses dans l’île. Ses gros fruits consommés par la population limitent sa dissémination" (Meyer, 2008; p. 27).
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Pagan Island	introduced	Ohba, T. (1994) (p. 58) Voucher cited: CBM-BS-58980
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Rota Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 80) Cultivated?
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Tinian Island	introduced	Raulerson, L. (2006) (p. 31)
Cook Islands Southern Cook Islands	Aitutaki Atoll	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Cook Islands Southern Cook Islands	‘Atiu Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Cook Islands Southern Cook Islands	Mangaia Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Cook Islands Southern Cook Islands	Ma‘uke Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Cook Islands Southern Cook Islands	Miti‘aro Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Cook Islands Southern Cook Islands	Rarotonga Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Ecuador (Galápagos Islands) Floreana Group	Floreana Island	introduced invasive cultivated	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Isabela Group	Isabela Island	introduced invasive cultivated	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Isabela Group	Volcán Sierra Negra, Isabela Island	introduced invasive cultivated	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) San Cristóbal Group	San Cristóbal Island	introduced invasive cultivated	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Cruz Group	Santa Cruz Island	introduced invasive cultivated	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santiago Group	Santiago Island	introduced invasive cultivated	Charles Darwin Foundation (2008)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 80) Cultivated?
Fiji Fiji Islands	Viti Levu Island	introduced cultivated	Smith, Albert C. (1981) (pp. 115-116)
French Polynesia Gambier Islands	Akamaru Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Gambier Islands	Aukena Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Gambier Islands	Taravai Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Fatu Hiva Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Fatu Hiva Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Tahuata (Tuhuata) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008)
French Polynesia Society Islands	Bora Bora Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Huahine Island	introduced invasive	Welsh, S. L. (1998) (pp. 132-133)
French Polynesia Society Islands	Huahine Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Maiao (Maiaoiti, Mauiti, Tupuaemanu) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Maupiti (Maurua) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Moorea Island	introduced invasive	Welsh, S. L. (1998) (pp. 132-133)
French Polynesia Society Islands	Moorea Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Welsh, S. L. (1998) (pp. 132-133)
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Taha‘a Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Tahiti Island	introduced invasive	Welsh, S. L. (1998) (pp. 132-133)
French Polynesia Society Islands	Tahiti Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Society Islands	Tetiaroa Atoll	introduced cultivated	Sachet, M.-H./Fosberg, F. R. (1983) (p. 64)
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Tubuai (Austral) Islands	Raivavae (Raevavae) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Tubuai (Austral) Islands	Rapa Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Tubuai (Austral) Islands	Rimatara (Rimitara) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Tubuai (Austral) Islands	Rurutu Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
French Polynesia Tubuai (Austral) Islands	Tubuai (Tupuai) Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
Guam Guam Island	Guam Island	introduced cultivated	Stone, Benjamin C. (1970) (p. 282) Vouchers cited: Stone 4121 (GUAM), Stone 4960 (GUAM)
Guam Guam Island	Guam Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 80)
State of Hawaii Hawaiian Islands	Hawai‘i (Big) Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 847-848)
State of Hawaii Hawaiian Islands	Kaua‘i Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 847-848)
State of Hawaii Hawaiian Islands	Maui Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 847-848)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive cultivated	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 847-848)
Marshall Islands Ratak Chain	Majuro (Mãjro) Atoll	introduced cultivated	Vander Velde, Nancy (2003) (p. 102)
Nauru Nauru Island	Nauru Island	introduced cultivated	Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (pp. 159-160)
New Caledonia Îles Loyauté (Loyalte Islands)	Île Maré	introduced cultivated	MacKee, H. S. (1994) (p. 72) Voucher cited: Baumann 14729 (Z)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced cultivated	MacKee, H. S. (1994) (p. 72) Vouchers cited: Baumann 5366 (Z), MacKee 13268
New Zealand (offshore islands) Kermadec Islands	Raoul Island	introduced cultivated	Sykes, W. R. (1977) (p. 111)
Niue Niue	Niue Island	introduced cultivated	Sykes, W. R. (1970) (p. 108) Voucher cited: CHR 168827
Niue Niue	Niue Island	introduced cultivated	Yuncker, T. G. (1943) (p. 55) Voucher cited: Yuncker 10087 (BISH)
Niue Niue	Niue Island	introduced cultivated	Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 45)
Palau Palau (main island group)	Babeldaob Island	introduced cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 102)
Palau Palau (main island group)	Babeldaob Island	introduced cultivated	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Palau Palau (main island group)	Koror Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 80)
Palau Palau (main island group)	Koror Island	introduced cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau Palau (main island group)	Koror Island		Beleu National Museum (Republic of Palau. Koror.) (1967) (voucher ID: BNM 412) Taxon name on voucher: Persea americana Mill.
Palau Palau (main island group)	Ngerkebesang Island	introduced cultivated	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Pitcairn Islands Pitcairn Islands	Pitcairn Island	introduced	Meyer, Jean-Yves (2000) (p. 104) "Potential invader; secndary forest".
Pitcairn Islands Pitcairn Islands	Pitcairn Island	introduced	St. John, H. (1987) (p. 34) Vouchers cited: Fosberg & R. Clark 11306, Ross 8
Pitcairn Islands Pitcairn Islands	Pitcairn Island	introduced invasive cultivated	Florence, J. (2004) (pp. 147-148)
Samoa Western Samoa Islands	Savai‘i Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Samoa Western Samoa Islands	Upolu Island	introduced cultivated	Space, James C./Flynn, Tim (2002) (p. Field notes.)
Solomon Islands Solomon Islands	Solomon Islands	introduced cultivated	Hancock, I. R./Henderson, C. P. (1988) (p. 98)
Tonga Ha‘apai Group	Lifuka and Foa Islands	introduced cultivated	Space, James C./Flynn, Tim (2001) (p. 74)
Tonga Tonga Islands	Tonga Islands	introduced cultivated	Yuncker, T. G. (1959) (p. 118) "Occasional in villages".
Tonga Tongatapu Group	Tongatapu Island	introduced cultivated	Space, James C./Flynn, Tim (2001) (p. 55)
United States (other Pacific offshore islands) Johnston Atoll	Johnston Island	introduced cultivated	Amerson, A. Binion, Jr./Shelton, Philip C. (1976) (p. 55)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Costa Rica Costa Rica	Costa Rica (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Honduras Honduras	Honduras (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Mexico Mexico	Mexico (United Mexican States)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Nicaragua Nicaragua	Nicaragua (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)
Panama Panama	Panama (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2009)

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia (Indian Ocean offshore islands) Christmas Island Group	Christmas Island	introduced cultivated	Swarbrick, J. T. (1997) (p. 123)
British Indian Ocean Territory Chagos Archipelago	Diego Garcia Island	introduced	Topp, J. M. W. (1988) (p. 13)
Seychelles Seychelles Islands	Agalega Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Stoddart, D. R. (1983) (p. 126)
Seychelles Seychelles Islands	Cousin Island	introduced cultivated	Fosberg, F. R. (1983) (p. 25) Voucher cited: Fosberg 52128 (US)
Seychelles Seychelles Islands	Frégate Island		Robertson, S. A./Todd, D. M. (1983) (p. 53)

Additional information:
Information from the Purdue University NewCROP web site.
Fact sheet from "Common forest trees of Hawaii" (PDF format).
Information from the World Agroforestry Centre's AgroForestryTree Database.

Additional online information about Persea americana is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Persea americana as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Persea americana may be available from the Germplasm Resources Information Network (GRIN).

References:

Amerson, A. Binion, Jr./Shelton, Philip C. 1976. The natural history of Johnston Atoll. Atoll Research Bulletin No. 192. Smithsonian Institution, Washington. 479 pp.

Beleu National Museum (Republic of Palau. Koror.). 1967. Voucher specimen #BNM412(Emmons, B. 37).

Charles Darwin Foundation. 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Charles Darwin Foundation, Galapagos, Ecuador.

Charles Darwin Research Station. 2005. CDRS Herbarium records.

Cronk, Q. C. B./Fuller, J. L. 2001. Plant invaders. Earthscan Publications, Ltd., London. 241 pp.

Florence, J. 2004. Flore de la Polynésie française, Vol. 2. Paris. IRE Editions, Publications Scientifiques, Collection Faune et Flore Tropicales 41. 503 pp.

Fosberg, F. R. 1983. Natural history of Cousin Island. In: Sachet, M. H., D. R. Stoddart, and F. R. Fosberg. Floristics and ecology of Western Indian Ocean islands. Atoll Research Bulletin No. 273. Smithsonian Institution, Washington. 253 pp.

Fosberg, F. R./Otobed, D./Sachet, M.-H./Oliver, R. L./Powell, D. A./Canfield, J. E. 1980. Vascular plants of Palau with vernacular names. Department of Botany, The Smithsonian Institution, Washington, D.C. 43 pp.

Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.

Fosberg, F. R./Sachet, Marie-Hélène/Stoddart, D. R. 1983. List of the recorded vascular flora of Agalega. In: Sachet, M. H., D. R. Stoddart, and F. R. Fosberg. Floristics and ecology of Western Indian Ocean islands. Atoll Research Bulletin No. 273. Smithsonian Institution, Washington. 253 pp.

Hancock, I. R./Henderson, C. P. 1988. Flora of the Solomon Islands. Research Bulletin No. 7. Ministry of Agriculture and Lands, Honiara. 203 pp.

Kostermans, André J. G. H. 1974. Lauracées. Flore de la Nouvelle-Calédonie et Dépendances. Muséum National d'Histoire Naturelle, Paris. Vol. 5, 123 pp.

Little, Elbert L./Skolmen, Roger G. 1989. Common forest trees of Hawaii (native and introduced). USDA Agriculture Handbook 679. Washington, D.C. 377 pp. + plates.

Lorence, David H./Wagner, Warren L. 2008. Flora of the Marquesas Islands. National Tropical Botanical Garden and the Smithsonian Institution. Online database.

MacKee, H. S. 1994. Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie. Muséum National d'Histoire Naturelle, Paris, 164 p.

McCormack, Gerald. 2008. Cook Islands Biodiversity Database, Version 2007.2. Cook Islands Natural Heritage Trust, Rarotonga.

McMullen, C. K. 1999. Flowering plants of the Galápagos. Comstock Pub. Assoc., Ithaca, N.Y. 370 p.

Meyer, Jean-Yves. 2000. Preliminary review of the invasive plants in the Pacific islands (SPREP Member Countries). In: Sherley, G. (tech. ed.). Invasive species in the Pacific: A technical review and draft regional strategy. South Pacific Regional Environment Programme, Samoa. 190 pp.

Meyer, Jean-Yves. 2008. Rapport de mission d'expertise a Rapa Nui du 02 au 11 Juin 2008: Plan d'action strategique pour lutter contre les plantes introduites envahissantes sur Rapa Nui (Île de pâques) [Strategic action plan to control invasive alien plants on Rapa Nui (Easter Island) [unpublished report]. Délégation à la Recherche, Ministère de l'Education, l'Enseignement supérieur et la Recherche, B.P. 20981 Papeete, Tahiti, Polynésie française. 62 pp. .

Ohba, T. 1994. Flora and vegetation of the northern Mariana Islands, Micronesia. Biological Expedition to the Northern Mariana Islands, Micronesia. In A. Asakua & T. Furuki, eds.  Natural History Research, Special issue no. 1, Chiba, Japan: Natural History Museum and Institute. pp. 13-69.

Raulerson, L. 2006. Checklist of Plants of the Mariana Islands. University of Guam Herbarium Contribution 40:1-69. .

Robertson, S. A./Todd, D. M. 1983. Vegetation of Frégate Island, Seychelles. In: Sachet, M. H., D. R. Stoddart, and F. R. Fosberg. Floristics and ecology of Western Indian Ocean islands. Atoll Research Bulletin No. 273. Smithsonian Institution, Washington. 253 pp.

Sachet, M.-H./Fosberg, F. R. 1983. An ecological reconnaissance of Tetiaroa Atoll, Society Islands. Atoll Research Bulletin No. 275. Smithsonian Institution, Washington. 67 pp. + photos.

Smith, Albert C. 1981. Flora Vitiensis nova: a new flora of Fiji. National Tropical Botanical Garden, Lawai, Kauai, Hawaii. Volume 2. 810 pp.

Soria, M./Taylor, U./Tye, A./Wilkenson, S. R. 2002. Identificación y manajo de malezas en las isles Galapágos. Fundación Charles Darwin, Parque Nacional Galapágos. 69 pp.

Space, James C./Flynn, Tim. 2001. Report to the Kingdom of Tonga on invasive plant species of environmental concern.  USDA Forest Service, Honolulu. 78 pp.

Space, James C./Flynn, Tim. 2002. Report to the Government of the Cook Islands on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 146 pp.

Space, James C./Flynn, Tim. 2002. Report to the Government of Samoa on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 83 pp.

Space, James C./Lorence, David H./LaRosa, Anne Marie. 2009. Report to the Republic of Palau: 2008 update on Invasive Plant Species. USDA Forest Service, Hilo, Hawai‘i. (In press).

Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate. 2004. Report to the Government of Niue and the United Nations Development Programme: Invasive plant species on Niue following Cyclone Heta. UNDP NIU/98/G31 - Niue Enabling Activity. 80 pp.

Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar. 2003. Report to the Republic of Palau on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 174 pp.

St. John, H. 1987. An account of the flora of Pitcairn Island with new Pandanus species. Honolulu, 65 pp.

Stone, Benjamin C. 1970. The flora of Guam. Micronesica 6:1-659.

Swarbrick, J. T. 1997. Environmental weeds and exotic plants on Christmas Island, Indian Ocean: a report to Parks Australia. 101 pp. plus appendix.

Sykes, W. R. 1970. Contributions to the flora of Niue. New Zealand Department of Scientific and Industrial Research Bulletin 200, Wellington. 321 pp.

Sykes, W. R. 1977. Kermadec Islands flora: an annotated checklist. New Zealand Department of Scientific and Industrial Research Bulletin 219, Wellington. 216 pp.

Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. 1994. The flora of Nauru. Smithsonian Institution, Washington. Atoll Research Bulletin 392:1-223.

Topp, J. M. W. 1988. An annotated check list of the flora of Diego Garcia, British Ocean Territory. Atoll Research Bulletin No. 313. Smithsonian Institution, Washington. 21 pp.

U.S. Dept. Agr., Agr. Res. Serv. 2009. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.

Vander Velde, Nancy. 2003. The vascular plants of Majuro Atoll, Republic of the Marshall Islands. Smithsonian Institution, Atoll Research Bulletin No. 503:1-141.

Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawai‘i Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).

Welsh, S. L. 1998. Flora Societensis: A summary revision of the flowering plants of the Society Islands. E.P.S. Inc., Orem, Utah. 420 pp.

Whistler, W. A. 1984. Annotated list of Samoan plant names. Economic Botany 38:464.489.

Whistler, W. A. 1990. Ethnobotany of the Cook Islands: The plants, their Maori names, and their uses. Allertonia 5(4):347-424.

Wiggins, I. L./Porter, D. M. 1971. Flora of the Galapágos Islands. Stanford University Press. 998 pp.

Yuncker, T. G. 1943. The flora of Niue Island. Bishop Mus. Bull. 178. Bishop Museum Press, Honolulu. 126 pp.

Yuncker, T. G. 1959. Plants of Tonga. Bishop Museum Bull. 220. Bishop Museum Press, Honolulu. 343 pp.