|
(Willd.) Zizka, Poaceae |
|
Present on Pacific Islands? yes
Threat only at high elevations? no
Other Latin names: Rhynchelytrum repens (Willd.) C.E. Hubbard, Tricholaena rosea Nees, Tricholaena repens (Willd.) Hitchc.
Common name(s): [more details]
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English: blanketgrass, Holme's grass, Natal grass, Natal redtop, red Natal grass |
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Fijian: thongithongi |
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French: fleur-la-misère, herbe du Natal, herbe pappangue, herbe rose, tit fleur, tricholène |
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Spanish: paja rosada, pasto Natal, zacate rosado |
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Tongan: salapona |
Habit: grass
Description: "Perennial [grass]; culms up to 90-100 cm high, in age tufted, glabrous, branched; sheaths to 10 cm long, sometimes longer, lower ones often slightly pubescent; ligule of short hairs 1 mm long; blades flat, 5-20 cm long, 2-5 mm wide, usually glaucous, glabrous dorsally; panicles to about 15 cm long, red or purplish at first, later silvery-pink, of fine ascending scabridulous branchlets; spikelets about 5 mm long, clothed with fine silky hairs 3-5 mm long (chiefly on upper glume and sterile lemma); caryopsis ovate, about 1.3 mm long, smooth, light brown" (Stone, 1970; p. 222).
Description from World Grass Species.
Habitat/ecology: In Hawaii, "naturalized in disturbed, usually dry areas, including urban and agricultural land, dry shrubland, dry forest, and subalpine forest, 0-1,950 m" (Wagner et al., 1999; p. 1588); "displaces low-growing natives; a fire hazard" (Motooka et al., 2003). In Fiji, "common from sea level to about 600 m along roadsides and in cultivated areas, on hillsides, and along forested streams in reed country" (Smith, 1979; p. 363). In New Caledonia, "introduit vers 1900 comme plante fourragère. Maintenant très commun et voyant à basse altitude en terrains vagues et bords de chemins" (MacKee, 1994; pp. 61-62).
Propagation: Seed
Native range: South Africa, now widely introduced.
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia (Pacific offshore islands)
Norfolk Islands |
Norfolk Island |
introduced
invasive |
Orchard, Anthony E., ed. (1994) (p. 12)
"An occasional weed". Vouchers cited: P. Ralston 50 (CHR), W.R. Sykes NI 605 (CHR) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 62)
as Tricholaena rosea Nees |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
Bishop Museum (Honolulu) (1950) (voucher ID: BISH 127193)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 62)
as Tricholaena rosea Nees |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
Bishop Museum (Honolulu) (1985) (voucher ID: BISH 572183)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Cook Islands
Southern Cook Islands |
Atiu Island | McCormack, Gerald (2007) | |
|
Cook Islands
Southern Cook Islands |
Atiu Island |
introduced
invasive |
Space, James C./Flynn, Tim (2002) (p. 81) |
|
Cook Islands
Southern Cook Islands |
Atiu Island |
Bishop Museum (Honolulu) (1991) (voucher ID: BISH 664900)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Cook Islands
Southern Cook Islands |
Atiu Island |
Bishop Museum (Honolulu) (1991) (voucher ID: BISH 664901)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Cook Islands
Southern Cook Islands |
Mangaia Island | McCormack, Gerald (2007) | |
|
Cook Islands
Southern Cook Islands |
Mangaia Island |
introduced
invasive |
Space, James C./Flynn, Tim (2002) (p. 87) |
|
Cook Islands
Southern Cook Islands |
Mangaia Island |
Bishop Museum (Honolulu) (1990) (voucher ID: BISH 614722)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Cook Islands
Southern Cook Islands |
Mauke Island | McCormack, Gerald (2007) | |
|
Cook Islands
Southern Cook Islands |
Mauke Island |
introduced
invasive |
Space, James C./Flynn, Tim (2002) (p. 93) |
|
Cook Islands
Southern Cook Islands |
Rarotonga Island | McCormack, Gerald (2007) | |
|
Cook Islands
Southern Cook Islands |
Rarotonga Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 666138)
Taxon name on voucher: Rhynchelytrum repens (Willd.) C.E.Hubb. |
|
|
Fiji
Fiji Islands |
Vanua Levu Island |
introduced
invasive cultivated |
Smith, Albert C. (1979) (p. 363)
Vouchers cited: DA 3669, DA 8810, DA 8743 |
|
Fiji
Fiji Islands |
Viti Levu Island |
introduced
invasive cultivated |
Smith, Albert C. (1979) (p. 363)
Vouchers cited: DA 8235, DA 8247, DA 8269, Smith 4259, DA 8285, DA 7972, DA 7930, DA 8107, DA 1648, DA 7480 |
|
Fiji
Fiji Islands |
Viti Levu Island |
Bishop Museum (Honolulu) (1954) (voucher ID: BISH 31603)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Fiji
Fiji Islands |
Viti Levu Island |
Bishop Museum (Honolulu) (1939) (voucher ID: BISH 31604)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Fiji
Fiji Islands |
Viti Levu Island |
Bishop Museum (Honolulu) (1947) (voucher ID: BISH 126677)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
Bishop Museum (Honolulu) (1987) (voucher ID: BISH 541055)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
Bishop Museum (Honolulu) (1984) (voucher ID: BISH 544959)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 558057)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Bora Bora Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Voucher cited: BRY 25436 |
|
French Polynesia
Society Islands |
Bora Bora Island |
Bishop Museum (Honolulu) (1993) (voucher ID: BISH 663576)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Huahine Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Voucher cited: Krauss 1660 |
|
French Polynesia
Society Islands |
Huahine Island |
Bishop Museum (Honolulu) (1979) (voucher ID: BISH 619019)
Taxon name on voucher: Rhynchelytrum repens (Willd.) C.E.Hubb. |
|
|
French Polynesia
Society Islands |
Huahine Island |
Bishop Museum (Honolulu) (1992) (voucher ID: BISH 643637)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Maupiti (Maurua) Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Voucher cited: Fosberg 64921 |
|
French Polynesia
Society Islands |
Maupiti (Maurua) Island |
Bishop Museum (Honolulu) (1985) (voucher ID: BISH 614448)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Voucher cited: Florence 8232 |
|
French Polynesia
Society Islands |
Moorea Island |
Bishop Museum (Honolulu) (1967) (voucher ID: BISH 22336)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Moorea Island |
Bishop Museum (Honolulu) (1983) (voucher ID: BISH 492567)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Moorea Island |
Bishop Museum (Honolulu) (1987) (voucher ID: BISH 547517)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Tahaa Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Voucher cited: Fosberg 63434 |
|
French Polynesia
Society Islands |
Tahaa Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 586744)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Tahaa Island |
Bishop Museum (Honolulu) (1990) (voucher ID: BISH 642428)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Vouchers cited: Florence 2286, BRY 26046 |
|
French Polynesia
Society Islands |
Tahiti Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491710)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Society Islands |
Tahiti Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 586752)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Tuamotu Archipelago |
Fangataufa Atoll |
Bishop Museum (Honolulu) (1987) (voucher ID: BISH 664032)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Tuamotu Archipelago |
Fangataufa Atoll |
Bishop Museum (Honolulu) (1987) (voucher ID: BISH 664036)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Tuamotu Archipelago |
Makatea (Maatea) Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 492122)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Tuamotu Archipelago |
Niau Atoll |
Bishop Museum (Honolulu) (1990) (voucher ID: BISH 642824)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
French Polynesia
Tuamotu Archipelago |
Tikehau (Tikahau) Atoll |
Bishop Museum (Honolulu) (1984) (voucher ID: BISH 546534)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Guam
Guam Island |
Guam Island |
introduced
invasive cultivated |
Stone, Benjamin C. (1970) (p. 222)
Vouchers cited: Stone 4371 (GUAM), Stone 4715 (GUAM) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 62)
as Tricholaena rosea Nees |
|
Guam
Guam Island |
Guam Island |
Bishop Museum (Honolulu) (1953) (voucher ID: BISH 127192)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Guam
Guam Island |
Guam Island |
Bishop Museum (Honolulu) (1962) (voucher ID: BISH 127194)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Guam
Guam Island |
Guam Island |
Bishop Museum (Honolulu) (1965) (voucher ID: BISH 588366)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
State of Hawaii
Hawaiian Islands |
Kahoolawe Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
State of Hawaii
Hawaiian Islands |
Lānai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
State of Hawaii
Hawaiian Islands |
Maui Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
State of Hawaii
Hawaiian Islands |
Molokai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
State of Hawaii
Hawaiian Islands |
Niihau Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
Japan (offshore islands)
Bonin (Ogasawara) Islands |
Bonin (Ogasawara) Islands |
introduced
|
Kato, Hidetoshi (2007)
As Rhynchelytrum repens (Willd.) C.E.Hubb. |
|
Kiribati
Line Islands |
Tabuaeran (Fanning) Atoll |
introduced
invasive |
Wester, Lyndon (1985) (p. 17) |
|
Kiribati
Tungaru (Gilbert) Islands |
Tarawa Atoll |
Swarbrick, John T. (1997) (p. 96)
Presence not verified. |
|
|
Marshall Islands
Ralik Chain |
Enewetak (Ãnewetak) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 62) |
|
Marshall Islands
Ralik Chain |
Enewetak (Ãnewetak) Atoll |
introduced
invasive |
St. John, Harold (1960) (p. 323)
Voucher cited: Taylor 46-1296 |
|
Nauru
Nauru Island |
Nauru Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 62) |
|
Nauru
Nauru Island |
Nauru Island |
introduced
invasive |
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 73)
As Tricholaena rosea Nees; vouchers cited: Fosberg 58655 (US), Scully 170N (US?), Thaman & Manner 49 (SUVA), Swarbrick 9567 |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Îles Loyauté (Loyalty Islands) |
introduced
invasive cultivated |
Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 381)
Rudérale très commune et voyante à basse altitude. |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Îles Loyauté (Loyalty Islands) |
introduced
invasive cultivated |
MacKee, H. S. (1994) (pp. 61-62) |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Maré |
introduced
invasive cultivated |
MacKee, H. S. (1994) (pp. 61-62)
Voucher cited: Däniker 2619 |
|
New Caledonia
New Caledonia Archipelago |
Île Belep |
introduced
invasive cultivated |
Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 381)
Rudérale très commune et voyante à basse altitude. |
|
New Caledonia
New Caledonia Archipelago |
Île Belep |
introduced
invasive cultivated |
MacKee, H. S. (1994) (pp. 61-62) |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive cultivated |
Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 381)
Rudérale très commune et voyante à basse altitude. |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive cultivated |
MacKee, H. S. (1994) (pp. 61-62) |
|
Philippines
Philippine Islands |
Philippine Islands | Hafliger, Ernst/Scholz, Hildemar (1980) (p. 118) | |
|
Solomon Islands
Solomon Islands |
Solomon Islands | Swarbrick, John T. (1997) (p. 96) | |
|
Solomon Islands
Solomon Islands |
Solomon Islands |
introduced
invasive |
Hancock, I. R./Henderson, C. P. (1988) (p. 105) |
|
Tonga
Haapai Group |
Kotu Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 640404)
Taxon name on voucher: Rhynchelytrum repens (Willd.) C.E.Hubb. |
|
|
Tonga
Tonga Islands |
Tonga Islands |
Yuncker, T. G. (1959) (p. 62)
"Occasional throughout the group". |
|
|
Tonga
Tongatapu Group |
Eua Island |
Yuncker, T. G. (1959) (p. 62)
Voucher cited: Yuncker 15490 |
|
|
Tonga
Tongatapu Group |
Eua Island |
Bishop Museum (Honolulu) (1953) (voucher ID: BISH 127186)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Whistler, W. A. (1992) (p. 101)
Vouchers cited: Setchell & Parks 15346, Soakai 68, Soakai 752 |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Yuncker, T. G. (1959) (p. 62)
Vouchers cited: Hürlmann 39, Yuncker 15151 |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Bishop Museum (Honolulu) (1953) (voucher ID: BISH 127188)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Bishop Museum (Honolulu) (1926) (voucher ID: BISH 127189)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
Tonga
Vavau Group |
Vavau Island |
Whistler, W. A. (1992) (p. 101)
Voucher cited: Soakai 873 |
|
|
Tonga
Vavau Group |
Vavau Island |
Yuncker, T. G. (1959) (p. 62)
Voucher cited: Yuncker 16068 |
|
|
Tonga
Vavau Group |
Vavau Island |
Bishop Museum (Honolulu) (1953) (voucher ID: BISH 127187)
Taxon name on voucher: Melinis repens (Willd.) Zizka |
|
|
United States (other Pacific offshore islands)
Midway Atoll |
Midway Atoll |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1588, 1912) |
|
Vanuatu
New Hebrides Islands |
Vanuatu (Republic of) | Swarbrick, John T. (1997) (p. 96) | |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis and Futuna Islands | Whistler, W. A. (1988) (p. 56) | |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia
Australia (continental) |
Australia (continental) | Hafliger, Ernst/Scholz, Hildemar (1980) (p. 118) | |
|
Cambodia
Cambodia |
Cambodia (Kingdom of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 307) | |
|
China
China |
China (People's Republic of) | Hafliger, Ernst/Scholz, Hildemar (1980) (p. 118) | |
|
Colombia
Colombia |
Colombia (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 307) | |
|
Honduras
Honduras |
Honduras (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 307) | |
|
Indonesia
Indonesia |
Indonesia (Republic of) | Hafliger, Ernst/Scholz, Hildemar (1980) (p. 118) | |
|
Malaysia
Malaysia |
Malaysia (country of) | Hafliger, Ernst/Scholz, Hildemar (1980) (p. plate) | |
|
Mexico
Mexico |
Mexico (United Mexican States) | Hafliger, Ernst/Scholz, Hildemar (1980) (p. 118) | |
|
Taiwan
Taiwan |
Taiwan Island | Hafliger, Ernst/Scholz, Hildemar (1980) (p. plate) | |
|
Thailand
Thailand |
Thailand (Kingdom of) | Hafliger, Ernst/Scholz, Hildemar (1980) (p. plate) | |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia (Indian Ocean offshore islands)
Christmas Island Group |
Christmas Island |
introduced
invasive |
Swarbrick, J. T. (1997) (p. 125) |
|
La Réunion (France)
La Réunion Island |
La Réunion Island |
introduced
invasive |
Lavergne, Christophe (2006)
"Envahissant" |
|
Mauritius
Mautitius Islands (Mauritius and Rodrigues) |
Mauritius Island | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 307) | |
|
Seychelles
Seychelles Islands |
Seychelles Islands |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2007) |
Control:
Chemical: "HAVO staff reported control with glyphosate at 1% of product sprayed to wet (Chris Zimmer, HAVO)" (Motooka et al., 2003).
Additional information: Information from the book "Weeds
of Hawaiis Pastures and Natural Areas; An Identification and Management Guide"
(Motooka et al., 2003). (PDF format).
Additional online information about Melinis repens is available from the Hawaiian Ecosystems at Risk project (HEAR).
Taxonomic information about Melinis repens may be available from the Germplasm Resources Information Network (GRIN).
References:
Bishop Museum (Honolulu). 1926. Voucher specimen #BISH 127189 (Setchell, W.A. 15346).
Bishop Museum (Honolulu). 1939. Voucher specimen #BISH 31604 (Parham, B.E. 1648).
Bishop Museum (Honolulu). 1947. Voucher specimen #BISH 126677 (Smith, A.C. 4259).
Bishop Museum (Honolulu). 1950. Voucher specimen #BISH 127193 (Fosberg, F.R. 31298).
Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 127186 (Yuncker, T.G. 15490).
Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 127187 (Yuncker, T.G. 16068).
Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 127188 (Yuncker, T.G. 15151).
Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 127192 (Fosberg, F.R. 35252).
Bishop Museum (Honolulu). 1954. Voucher specimen #BISH 31603 (Koroiveibau, D. 7480).
Bishop Museum (Honolulu). 1962. Voucher specimen #BISH 127194 (Stone, B.C.M. 4471).
Bishop Museum (Honolulu). 1965. Voucher specimen #BISH 588366 (Evans, M. 222).
Bishop Museum (Honolulu). 1967. Voucher specimen #BISH 22336 (Smith, H.M. 29).
Bishop Museum (Honolulu). 1979. Voucher specimen #BISH 619019 (N.Krauss 1660).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491710 (Florence, J. 2286).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 492122 (Florence, J. 3080).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 558057 (Florence, J. 4174).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 586744 (Fosberg, F.R. 63434).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 586752 (Fosberg, F.R. 63743).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 640404 (G.Buelow 3227).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 666138 (Sykes, W.R. 399911).
Bishop Museum (Honolulu). 1983. Voucher specimen #BISH 492567 (Florence, J. 4717).
Bishop Museum (Honolulu). 1984. Voucher specimen #BISH 544959 (Florence, J. 6948).
Bishop Museum (Honolulu). 1984. Voucher specimen #BISH 546534 (Florence, J. 7105).
Bishop Museum (Honolulu). 1985. Voucher specimen #BISH 572183 (Raulerson, L. 10114).
Bishop Museum (Honolulu). 1985. Voucher specimen #BISH 614448 (Fosberg, F.R. 64921).
Bishop Museum (Honolulu). 1987. Voucher specimen #BISH 541055 (Florence, J. 8447).
Bishop Museum (Honolulu). 1987. Voucher specimen #BISH 547517 (Florence, J. 8232).
Bishop Museum (Honolulu). 1987. Voucher specimen #BISH 664032 (Jolinon, J.C. 1311).
Bishop Museum (Honolulu). 1987. Voucher specimen #BISH 664036 (Jolinon, J.C. 1270).
Bishop Museum (Honolulu). 1990. Voucher specimen #BISH 614722 (Sykes, W.R. 467822).
Bishop Museum (Honolulu). 1990. Voucher specimen #BISH 642428 (Florence 10607).
Bishop Museum (Honolulu). 1990. Voucher specimen #BISH 642824 (Florence 10149).
Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 664900 (Sykes, W.R. 474751).
Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 664901 (Sykes, W.R. 474751).
Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 643637 (Florence 11702).
Bishop Museum (Honolulu). 1993. Voucher specimen #BISH 663576 (Florence 12069).
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Orchard, Anthony E., ed. 1994. Flora of Australia. Vol. 49, Oceanic islands 1. Australian Government Publishing Service, Canberra.
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Space, James C./Flynn, Tim. 2002. Report to the Government of the Cook Islands on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 146 pp.
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Swarbrick, J. T. 1997. Environmental weeds and exotic plants on Christmas Island, Indian Ocean: a report to Parks Australia. 101 pp. plus appendix.
Swarbrick, John T. 1997. Weeds of the Pacific Islands. Technical paper no. 209. South Pacific Commission, Noumea, New Caledonia. 124 pp.
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U.S. Dept. Agr., Agr. Res. Serv. 2007. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawaii Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).
Waterhouse, B. M./Mitchell, A. A. 1998. Northern Australia Quarantine Strategy: weeds target list. Second edition. Australian Quarantine & Inspection Service, Miscellaneous Publication No. 6/98. 110 pp.
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Whistler, W. A. 1988. Checklist of the weed flora of western Polynesia. Technical Paper No. 194, South Pacific Commission, Noumea, New Caledonia. 69 pp.
Whistler, W. A. 1992. Flora of Tonga [unpublished checklist]. 107 pp.
Yuncker, T. G. 1959. Plants of Tonga. Bishop Museum Bull. 220. Bishop Museum Press, Honolulu. 343 pp.