|
P.Beauv., Poaceae |
|
Present on Pacific Islands? yes
Primarily a threat at high elevations? no
Risk assessment results:
Reject, score: 7 (Go to the risk assessment (Australia)).
High risk, score: 18 (Go to the risk assessment (Pacific)).
Other Latin names: Panicum melinus Trin.; Panicum minutiflora (Beauv.) Rasp.
Common name(s): [more details]
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Chinese: tang mi cao |
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English: Brazilian stink grass, efwatakala grass, melinis, molasses grass, stink grass, wynne grass |
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French: herbe à miel, herbe à molasse, herbe molasse, herbe molasses, mélinis |
|
Rapa Nui: mauku piro |
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Spanish: chopín, pasto de gordura, pasto hediondo, pasto miel |
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Tongan: puakatau |
Habit: grass
Description: "Branching perennial [grass], the culms to 90 cm long, ascending; foliage viscid (sticky), fragrant, with odor like molasses; blades up to nearly 1 ft. long, often reddish, the ligule a row of hairs; panicles 10-20 cm long, slender, purplish when young; spikelets less than 2.5 mm long; sterile lemmas awned, the awn 12 mm long or less" (Stone, 1970; p. 200).
"Herbacée pérenne en touffe dense, atteignant 1 à 2 m de hauteur, reconnaissable à ses tiges et feuilles abondamment poilues et collantes à forte odeur d’huile, et ses inflorescences dressées rousses ou rougêatres" (Meyer, 2008; p. 20).
Habitat/ecology: "This spreading, perennial mat grass smothers everything around it. Once established, it forms monotypic stands from rooted runners. It is considered a good forage grass and therefore is not a candidate for biological control. It is adapted to fire, and the dense mats are generally only partly consumed. Regeneration from the remaining portions is rapid, and colony expansion into adjacent burned areas generally follows. It is found from sea level to 1,500 m in dry and mesic environments" in Hawaii (Smith, 1985; p. 195), to 2000 m in French Polynesia. In Hawaii, "naturalized and common in primarily dry to mesic, disturbed, usually open areas, 120-1,220 m" (Wagner et al., 1999; p. 1562). "A serious pest in dryer environments because of sprawling growth and fire hazard in dry seasons" (Motooka et al., 2003).
"Espèce pionnière, colonisant les zones ouvertes et incendiées (il s’agit d’une plante pyrophile ou « pyrophyte », c’est-à-dire tolérante et adaptée aux incendies), elle n’est relativement pas consommée par les animaux (Cronk & Fuller, 1995)" (Meyer, 2008; p. 20). In New Caledonia, "occupe en peuplements denses monospécifiques de grandes étendues de terrain. Peu commun aux Loyauté mais a été récolté à Tiga" (MacKee, 1994; p. 60).
Propagation: The seeds are dispersed by wind.
Native range: Africa; introduced to many locations.
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
American Samoa
Tutuila Islands |
Tutuila Island |
Bishop Museum (Honolulu) (1985) (voucher ID: BISH 495534)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
Australia (Pacific offshore islands)
Lord Howe Island |
Lord Howe Island |
introduced
|
Pickard, J. (1984) (p. 205) |
|
Australia (Pacific offshore islands)
Lord Howe Island |
Lord Howe Island |
introduced
invasive |
Orchard, Anthony E., ed. (1994) (p. 22)
"Introduced, probably relatively recently, as a fodder grass". Voucher cited: J. Pickard 3531 (NSW) |
|
Australia (Pacific offshore islands)
Norfolk Islands |
Norfolk Island |
introduced
invasive |
Orchard, Anthony E., ed. (1994) (p. 12)
"Introduced, probably relatively recently, as a fodder grass". Voucher cited: W.R. Sykes NI 547 (CHR) |
|
Chile (offshore islands)
Rapa Nui (Easter) Island |
Rapa Nui (Easter) Island (Isla de Pasqua) |
introduced
invasive |
Meyer, Jean-Yves (2008) (pp. 20, 29)
"Lors de cette mission, nous avons observé des tapis denses monospécifiques sur la majeure partie de l’île (excepté sur le sommet de Maunga Terevaka), notamment autour de Rano Kau, Rano Aroi et Rano Raraku où l’espèce menace directement les sites archéologiques. Il s’agit selon notre expérience de l’invasion la plus sévère et la plus spectaculaire du mélinis dans une île du Pacifique" (Meyer, 2008; p. 20-21). |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
introduced
|
Raulerson, L. (2006) (p. 10) |
|
Ecuador (Galápagos Islands)
San Cristóbal Group |
San Cristóbal Island |
introduced
invasive cultivated |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Santa Cruz Group |
Santa Cruz Island |
introduced
invasive cultivated |
Charles Darwin Foundation (2008) |
|
Fiji
Fiji Islands |
Viti Levu Island |
introduced
invasive cultivated |
Smith, Albert C. (1979) (p. 342)
Vouchers cited: DA 14349, DA 8318, DA 3525, DA 3526, DA 3615, DA 3617, DA 3659, DA 8009, DA 9170, DA 10139 |
|
Fiji
Fiji Islands |
Viti Levu Island |
Bishop Museum (Honolulu) (1965) (voucher ID: BISH 31713)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Vouchers cited: J. Florence, D.H. Lorence & W.L. Wagner 9500 (PAP), D.H. Lorence, L. Dunn, S.P. Perlman, K.R. Wood & J.-Y. Meyer 9023 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
Bishop Museum (Honolulu) (1988) (voucher ID: BISH 601874)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
Bishop Museum (Honolulu) (1988) (voucher ID: BISH 1000693)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1988) (voucher ID: PTBG 8517)
Taxon name on voucher: Melinis minutiflora |
|
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 21968)
Taxon name on voucher: Melinis minutiflora P. Beauv. |
|
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39296)
Taxon name on voucher: Melinis minutiflora |
|
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39560)
Taxon name on voucher: Melinis minutiflora |
|
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 6891 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
Bishop Museum (Honolulu) (1984) (voucher ID: BISH 554197)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: S.P. Perlman, K.R. Wood & J.-Y. Meyer 15866 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Society Islands |
Huahine Island |
introduced
|
Fosberg, F. R. (1997) (p. 102) |
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Vouchers cited: Cartier 143, Cartier 243 |
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Vouchers cited: C.L. Cartier 143 (PAP), C.L. Cartier 243 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
|
Fosberg, F. R. (1997) (p. 102) |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 339)
Voucher cited: Fosberg 61265 |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Vouchers cited: J. Florence 3193 (PAP), J. Florence 3276 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Society Islands |
Tahiti Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 491721)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Society Islands |
Tahiti Island |
Bishop Museum (Honolulu) (1982) (voucher ID: BISH 558030)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Society Islands |
Tahiti Island |
Bishop Museum (Honolulu) (1988) (voucher ID: BISH 582489)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Society Islands |
Tahiti Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1981) (voucher ID: PTBG 38249)
Taxon name on voucher: Melinis minutiflora P. Beauv. |
|
|
French Polynesia
Tuamotu Archipelago |
Niau Atoll |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 10079 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Tuamotu Archipelago |
Niau Atoll |
Bishop Museum (Honolulu) (1992) (voucher ID: BISH 642460)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Austral (Tubuai) Islands |
Raivavae (Raevavae) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Vouchers cited: J. Florence & W.R. Sykes 11200 (PAP), J. Florence & W.R. Sykes 11408 (PAP) Adventice, menace pour la biodiversité. |
|
French Polynesia
Austral (Tubuai) Islands |
Raivavae (Raevavae) Island |
Bishop Museum (Honolulu) (1992) (voucher ID: BISH 643773)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Austral (Tubuai) Islands |
Raivavae (Raevavae) Island |
Bishop Museum (Honolulu) (1992) (voucher ID: BISH 644107)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
French Polynesia
Austral (Tubuai) Islands |
Rurutu Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 5544 (PAP) Adventice, menace pour la biodiversité. |
|
Guam
Guam Island |
Guam Island |
introduced
cultivated |
Stone, Benjamin C. (1970) (pp. 200-201) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 49) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Meyer, Jean-Yves (2000) (p. 95)
"Potential invader". |
|
State of Hawaii
Hawaiian Islands |
Hawaii (Big) Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
State of Hawaii
Hawaiian Islands |
Kahoolawe Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
State of Hawaii
Hawaiian Islands |
Lānai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
State of Hawaii
Hawaiian Islands |
Maui Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
State of Hawaii
Hawaiian Islands |
Molokai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1562) |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Îles Loyauté (Loyalty Islands) |
introduced
invasive |
MacKee, H. S. (1994) (p. 60) |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Tiga |
introduced
invasive |
MacKee, H. S. (1994) (p. 60) |
|
New Caledonia
New Caledonia |
New Caledonia Islands |
introduced
invasive |
Meyer, Jean-Yves (2000) (p. 100)
"Fernlands" |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive cultivated |
Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 381)
Abondante, surtout sur schistes dans le Sud; occupe en peuplements denses monospécifiques de grandes étendues de terrain. |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive |
MacKee, H. S. (1994) (p. 60)
Voucher cited: MacKee 12716 |
|
Niue
Niue |
Niue Island |
introduced
|
Sykes, W. R. (1970) (pp. 244, 246)
Voucher cited: CHR 168746 |
|
Niue
Niue |
Niue Island |
introduced
|
Whistler, W. A. (1988) (p. 53) |
|
Niue
Niue |
Niue Island |
introduced
invasive |
Space, James C./Flynn, Tim (2000) (p. 5)
Voucher: Flynn 6711 (PTBG, US, NIUE) |
|
Niue
Niue |
Niue Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 12) |
|
Niue
Niue |
Niue Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2000) (voucher ID: PTBG 30202)
Taxon name on voucher: Melinis minutiflora Beauv. |
|
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 49) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 16) |
|
Palau
Palau (main island group) |
Babeldaob Island |
Beleu National Museum (Republic of Palau. Koror.) (1981) (voucher ID: BNM 3075)
Taxon name on voucher: Melinis minutiflora Beauv. |
|
|
Palau
Palau (main island group) |
Babeldaob Island |
Beleu National Museum (Republic of Palau. Koror.) (1981) (voucher ID: BNM 3140)
Taxon name on voucher: Melinis minutiflora Beauv. |
|
|
Papua New Guinea
Papua New Guinea (eastern New Guinea Island) |
Papua New Guinea (eastern New Guinea Island) |
introduced
invasive |
Henty, E. E. (1969) (p. 124)
Naturalized in numerous localities; an aggressive colonizer of recently cleared land. |
|
Solomon Islands
Solomon Islands |
Solomon Islands | Swarbrick, John T. (1997) (p. 94) | |
|
Solomon Islands
Solomon Islands |
Solomon Islands |
introduced
|
Hancock, I. R./Henderson, C. P. (1988) (p. 90) |
|
Tonga
Tongatapu Group |
Tongatapu Island |
introduced
invasive |
Space, James C./Flynn, Tim (2001) (p. 7)
Rare. |
|
Tonga
Tongatapu Group |
Tongatapu Island |
Whistler, W. A. (1992) (p. 100)
Vouchers cited: Soakai 244, Soakai 700 |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
introduced
invasive |
Yuncker, T. G. (1959) (p. 56)
Uncommon. Voucher cited: Yuncker 16229 |
|
Tonga
Tongatapu Group |
Tongatapu Island |
Bishop Museum (Honolulu) (1953) (voucher ID: BISH 125541)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Bishop Museum (Honolulu) (1960) (voucher ID: BISH 125542)
Taxon name on voucher: Melinis minutiflora P.Beauv. |
|
|
Tonga
Vavau Group |
Vavau Island |
introduced
invasive |
Space, James C./Flynn, Tim (2001) (p. 7) |
|
Vanuatu
New Hebrides Islands |
Vanuatu (Republic of) | Swarbrick, John T. (1997) (p. 94) | |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis (Uvea) Island |
introduced
invasive |
Orapa, Warea (2005)
Only known to be present on Wallis. |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis (Uvea) Island |
introduced
invasive cultivated |
Meyer, Jean-Yves (2007) (pp. 12, 27)
Localement naturalisée en bordure de piste et en zone à toafa près de la station fourragère expérimentale de Hihifo ainsi qu’au sommet du mont Lulu en bordure de route en zone à toafa. |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis (Uvea) Island |
introduced
invasive cultivated |
Meyer, Jean-Yves/Jourdan, Hervé/Malau, Atoloto (2008) (p. 9) |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis and Futuna Islands |
introduced
invasive |
Meyer, Jean-Yves (2000) (p. 107)
Fernlands |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia
Australia (continental) |
Australia (continental) |
introduced
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Australia
Australia (continental) |
Queensland |
introduced
invasive |
Queensland Herbarium (2002) (p. 5) |
|
China
China |
China (People's Republic of) |
introduced
invasive cultivated |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
Introduced into S. China as a fodder grass, locally naturalized. Hong Kong, Yunnan. |
|
China
China |
Hong Kong |
introduced
cultivated |
Wu, Te-lin (2001) (p. 340) |
|
Colombia
Colombia |
Colombia (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 233) | |
|
Costa Rica
Costa Rica |
Costa Rica (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 233) | |
|
Japan
Japan |
Japan (country) |
introduced
|
Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 192) |
|
Taiwan
Taiwan |
Taiwan Island |
introduced
invasive cultivated |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011) |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Comoros
Comoro Islands |
Comoro Islands |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
La Réunion (France)
La Réunion Island |
La Réunion Island |
introduced
invasive |
Lavergne, Christophe (2006)
"Envahissant" |
| Also reported from | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
United States (continental except west coast)
United States (other states) |
USA (Florida) |
introduced
|
U.S. Dept. Agr., Nat. Res. Cons. Serv. (2011) |
Comments: Because of its oil content, this grass can be a serious fire hazard. A serious invader in Hawaii, French Polynesia and Wallis and Futuna (Wagner et al., 1999, p. 1562; Meyer, 2000, p. 107).
Introduced into Guam, but not widely established (Stone, 1970; p. 200).
Control:
Chemical: "Sensitive to glyphosate. HAVO staff reported control with foliar application of glyphosate at 1% product in water (Chris Zimmer, HAVO)" (Motooka et al., 2003).
Additional information:
Excerpt from the book "Weeds
of Hawaiis Pastures and Natural Areas; An Identification and Management Guide"
(Motooka et al., 2003). (PDF format).
Information from the Purdue University NewCROP web site.
Fact sheet from the Government of Queensland, Australia. (PDF format).
Additional online information about Melinis minutiflora is available from the Hawaiian Ecosystems at Risk project (HEAR).
Information about Melinis minutiflora as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).
Taxonomic information about Melinis minutiflora may be available from the Germplasm Resources Information Network (GRIN).
References:
Beleu National Museum (Republic of Palau. Koror.). 1981. Voucher specimen #BNM3075(Timberlake, J. 3075).
Beleu National Museum (Republic of Palau. Koror.). 1981. Voucher specimen #BNM3140(Timberlake, J. 3140).
Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 125541 (Yuncker, T.G. 16229).
Bishop Museum (Honolulu). 1960. Voucher specimen #BISH 125542 (Hotta, M. 5151).
Bishop Museum (Honolulu). 1965. Voucher specimen #BISH 31713 (Qoro, I. 14349).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 491721 (Florence, J. 3276).
Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 558030 (Florence, J. 3193).
Bishop Museum (Honolulu). 1984. Voucher specimen #BISH 554197 (Florence, J. 6891).
Bishop Museum (Honolulu). 1985. Voucher specimen #BISH 495534 (Whistler, W.A. 5729).
Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 582489 (Gagne; collector's voucher number unknown).
Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 601874 (J.Florence 9500).
Bishop Museum (Honolulu). 1988. Voucher specimen #BISH 1000693 (W.L.Wagner 6167).
Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 642460 (Florence 10079).
Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 643773 (Florence 11408).
Bishop Museum (Honolulu). 1992. Voucher specimen #BISH 644107 (Florence 11200).
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