|
(L.) Lam., Convolvulaceae |
|
Present on Pacific Islands? yes
Threat only at high elevations? no
Common name(s): [more details]
|
Chamorro: kamote |
|
Chuukese: kómwu, kómwuti, kómwuutiy, omwuti, sachúmayimwo, yomwuutiy |
|
English: sweet potato |
|
Fijian: kawai ni vavalangi, kumala, kumara, ndambithi, wa uvi |
|
French: patate douce |
|
Hawaiian: uala, wala |
|
I-Kiribati: te kumara |
|
Kwaraae: butete |
|
Maori (Cook Islands): kuara, kūara, kūmala, kūmara, kumara |
|
Marshallese: biteto, piteto |
|
Niuean: kumara, simala, timala |
|
Other: gumala, kumara, oumera, tek |
|
Palauan: emutii |
|
Pukapukan: kūmala |
|
Samoan: umala |
|
Satawalese: kamote, komote |
|
Spanish: camote |
|
Tahitian: umara, umara |
|
Tongan: kumala, kumara |
|
Ulithian: komoti |
|
Woleaian: gamwuutiy |
|
Yapese: gamuti, kamuut |
Habit: vine
Description: "Creeping vine with milky sap, glabrous or pubescent, with enlarged tuberous edible roots; leaves ovate-orbicular, entire or palmately 3-7-lobed or -parted, cordate or subcordate, 4-15 cm long, 3-11 cm wide, on petioles commonly 3-15 cm long; flowers several to many in long-peduncled cymes; calyx-lobes briefly mucronate, about 10-15 mm long; corolla usually pale rose-violet, rarely albino, 3-5 cm long; darker (purplish) in the throat; ovary mostly pubescent; fruit a capsule" (Stone, 1970; p. 491).
Habitat/ecology: In Hawaii, "a Polynesian introduction, escaping cultivation and persisting near abandoned homesites and dumps" (Wagner et al., 1999; p. 555). In Fiji, "cultivated at elevations from near sea level to about 800 m and sometimes naturalized (up to 1,050 m) near cultivated areas, along roadsides, and in thickets along rivers and streams" (Smith, 1991; pp. 66-67). In New Caledonia, "on la rencontre en forêt ou dans des fourré secondaires près du littoral, sur schistes ou calcaires" (Heine, 1984; pp. 57-58).
Propagation: Seed
Native range: Cultivated worldwide, widely naturalized in tropics, origin neotropics but native range obscure (GRIN).
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Agrigan (Agrihan) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Agrigan (Agrihan) Island |
introduced
cultivated |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 37) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Alamagan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Alamagan Island |
introduced
cultivated |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 37) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Pagan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Pagan Island |
introduced
cultivated |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 37)
Vouchers cited: Fosberg 31357 (US), Anderson 595 (US, BISH,Fo), Anderson 596 (US, BISH, Fo) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
introduced
cultivated |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 37)
Voucher cited: Evans 2422 (US, BISH, Fo, NY) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
Cook Islands
Northern Cook Islands |
Manihiki Atoll |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Northern Cook Islands |
Nassau Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Northern Cook Islands |
Pukapuka Atoll |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Northern Cook Islands |
Rakahanga Atoll |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Northern Cook Islands |
Suwarrow Atoll |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Southern Cook Islands |
Atiu Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Southern Cook Islands |
Mangaia Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Southern Cook Islands |
Mauke Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Southern Cook Islands |
Mitiaro Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Southern Cook Islands |
Palmerston (Avarau) Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Cook Islands
Southern Cook Islands |
Rarotonga Island |
introduced
cultivated abo. intro. |
McCormack, Gerald (2007) |
|
Ecuador (Galápagos Islands)
Isabela Group |
Volcan Sierra Negra |
introduced
invasive cultivated |
Charles Darwin Research Station (2005) |
|
Ecuador (Galápagos Islands)
Santa Cruz Group |
Santa Cruz Island |
introduced
invasive cultivated |
Charles Darwin Research Station (2005) |
|
Federated States of Micronesia
Chuuk Islands |
Fefan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Chuuk Islands |
Murilo (Mwiriné) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Chuuk Islands |
Namoluk (Nómwunuuk) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Chuuk Islands |
Nomwin (Nómwiin) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Chuuk Islands |
Puluwat (Pwonowót) Atoll |
introduced
cultivated |
Manner, H. I./Mallon, E. (1989) (p. 56)
Voucher cited: Manner & Mallon 254 (GUAM), Manner & Mallon 260 (GUAM) |
|
Federated States of Micronesia
Chuuk Islands |
Satawan Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
invasive cultivated |
Glassman, S. F. (1952) (p. 99) |
|
Federated States of Micronesia
Yap Islands |
Eauripik Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Fais Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Fais Island |
introduced
cultivated |
Fosberg, F. R./Evans, M. (1969) (p. 12)
Voucher cited: Fosberg 46662 (US) |
|
Federated States of Micronesia
Yap Islands |
Faraulep (Faraulap) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Ifalik (Ifaluk) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Lamotrek Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Satawal Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Satawal Island |
introduced
cultivated |
Fosberg, F. R. (1969) (p. 10)
Voucher cited: Fosberg 46866 (US) |
|
Federated States of Micronesia
Yap Islands |
Ulithi Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Woleai Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Federated States of Micronesia
Yap Islands |
Yap (Waqab) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Fiji
Fiji Islands |
Lakemba Island |
introduced
invasive cultivated abo. intro. |
Smith, Albert C. (1991) (p. 67)
Voucher cited: Garnock-Jones 882 |
|
Fiji
Fiji Islands |
Viti Levu Island |
introduced
invasive cultivated abo. intro. |
Smith, Albert C. (1991) (p. 67)
Vouchers cited: Greenwood 256; Smith 4057, Smith 4740, Smith 9077, DA 1208, DA 2714, DA 6092 |
|
French Polynesia
Society Islands |
Maupiti (Maurua) Island |
introduced
cultivated abo. intro. |
Fosberg, F. R./Sachet, M.-H. (1987) (p. 58)
Vouchers cited: Morris 34 (US), Fosberg 64852 (US), Fosberg & Sachet 64967 (US) |
|
French Polynesia
Society Islands |
Moorea Island | University of California (2006) | |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
invasive abo. intro. |
Welsh, S. L. (1998) (pp. 84-85)
Vouchers cited: Moore 369, Moore 370, Moore 432, BRY 25731 |
|
French Polynesia
Society Islands |
Tahiti Island | Fosberg, F. R. (1997) (p. 31) | |
|
French Polynesia
Society Islands |
Tetiaroa Atoll |
introduced
cultivated |
Sachet, M.-H./Fosberg, F. R. (1983) (p. 65) |
|
French Polynesia
Tuamotu Archipelago |
Manihi Atoll |
introduced
|
Sachet, M-H. (1983) (p. 34) |
|
French Polynesia
Tuamotu Archipelago |
Raroia Atoll |
introduced
cultivated |
Doty, Maxwell S. (1954) (p. 39) |
|
French Polynesia
Tuamotu Archipelago |
Takapoto Atoll |
introduced
cultivated |
Sachet, M-H. (1983) (p. 34) |
|
Guam
Guam Island |
Guam Island |
introduced
abo. intro. |
Stone, Benjamin C. (1970) (p. 491) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 222) |
|
State of Hawaii
Hawaiian Islands |
Hawaii (Big) Island |
introduced
invasive cultivated abo. intro. |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 555) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive cultivated abo. intro. |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 555) |
|
State of Hawaii
Hawaiian Islands |
Maui Island |
introduced
invasive cultivated abo. intro. |
Oppenheimer, Hank L. (2003) (p. 10)
West Maui. Oppenheimer H79806 (BISH), Oppenheimer H10130 (BISH, PTBG) Escaping and persisting from cultivation. |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive cultivated abo. intro. |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 555) |
|
Kiribati
Line Islands |
Caroline Atoll |
introduced
cultivated |
Kepler, Angela K./Kepler, Cameron B. (1994) (p. 36) |
|
Kiribati
Line Islands |
Teraina (Washington) Island |
introduced
invasive cultivated |
Wester, Lyndon (1985) (p. 29) |
|
Kiribati
Tungaru (Gilbert) Islands |
Tarawa Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ralik Chain |
Ailinglaplap (Ailinglapalap) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ralik Chain |
Jaluit (Jãlwõj) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ralik Chain |
Jaluit (Jãlwõj) Atoll |
introduced
cultivated |
Fosberg, F. R./Sachet, M.-H. (1962) (p. 31)
Voucher cited: Fosberg 26748 (US) |
|
Marshall Islands
Ralik Chain |
Kili (Kõle) Atoll |
introduced
cultivated |
Vander Velde, Nancy/Vander Velde, Brian (2006) (p. 7)
"Rare; few plants in Bikini Projects garden". |
|
Marshall Islands
Ralik Chain |
Kwajalein (Kuwajleen) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ratak Chain |
Arno Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ratak Chain |
Arno Atoll |
Anderson, Donald (1951) (p. iii)
Voucher cited: Anderson 3784 |
|
|
Marshall Islands
Ratak Chain |
Arno Atoll |
introduced
cultivated |
Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 7)
"Uncommon; planted in Reimers' garden". |
|
Marshall Islands
Ratak Chain |
Likiep Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ratak Chain |
Likiep Atoll |
introduced
invasive cultivated |
Fosberg, F. R. (1955) (p. 17)
Voucher cited: Fosberg 27038 (US) Persisting from cultivation. |
|
Marshall Islands
Ratak Chain |
Majuro (Mãjro) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Marshall Islands
Ratak Chain |
Majuro (Mãjro) Atoll |
introduced
cultivated |
Vander Velde, Nancy (2003) (p. 84) |
|
Nauru
Nauru Island |
Nauru Island |
introduced
cultivated |
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 117) |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Maré |
introduced
invasive cultivated abo. intro. |
MacKee, H. S. (1994) (pp. 40-41)
Voucher cited: Baumann 14592 Spontané |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Îles Ouvéa (Ouvea Atoll) |
introduced
cultivated abo. intro. |
MacKee, H. S. (1994) (pp. 40-41)
Vouchers cited: Däniker 2372, Pradee in Veillon 1239 (NOU) |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive cultivated abo. intro. |
MacKee, H. S. (1994) (pp. 40-41)
Vouchers cited: Vieillard 1009, Pancher s.n., MacKee 17611, MacKee 20835, MacKee 22575, MacKee 33901 Spontané |
|
New Caledonia
New Caledonia Archipelago |
Île des Pins (Isle of Pines) |
introduced
invasive cultivated abo. intro. |
MacKee, H. S. (1994) (pp. 40-41)
Voucher cited: Baumann 13662 Spontané |
|
New Caledonia
New Caledonia Archipelago |
Île Ouen |
introduced
cultivated abo. intro. |
MacKee, H. S. (1994) (pp. 40-41)
Voucher cited: Bourret 2293 (NOU) |
|
New Zealand (offshore islands)
Kermadec Islands |
Raoul Island |
introduced
invasive |
Sykes, W. R. (1977) (p. 98) |
|
New Zealand (offshore islands)
Kermadec Islands |
Raoul Island |
introduced
invasive cultivated |
Webb, C. J./Sykes, W. R./Garnock-Jones, P. J. (1988) (p. 558)
"An occasional escape from cultivation". |
|
Niue
Niue |
Niue Island |
introduced
abo. intro. |
Sykes, W. R. (1970) (p. 72)
Vouchers cited: CHR 169785, CHR 169793 |
|
Niue
Niue |
Niue Island |
introduced
cultivated |
Yuncker, T. G. (1943) (pp. 98-99)
Voucher cited: Yuncker 9667 (BISH) |
|
Palau
Palau (main island group) |
Angaur Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 223) |
|
Pitcairn Islands
Pitcairn Islands |
Pitcairn Island |
introduced
|
St. John, H. (1987) (p. 47)
Voucher cited: Fosberg & R. Clark 11325 |
|
Solomon Islands
Solomon Islands |
Solomon Islands |
introduced
cultivated |
Whitmore, T. C. (1969) (p. 264) |
|
Solomon Islands
Solomon Islands |
Solomon Islands |
introduced
invasive cultivated |
Hancock, I. R./Henderson, C. P. (1988) (p. 83) |
|
Tonga
Tonga Islands |
Tonga Islands |
introduced
cultivated |
Yuncker, T. G. (1959) (p. 225)
"Cultivated throughout Tonga". |
|
Tonga
Tongatapu Group |
Tongatapu Island |
Ellison, Joanna C. (1990) (p. 28)
On outlier islet of Ata. |
|
|
Tuvalu
Tuvalu (Ellis) Islands |
Nui Atoll |
Woodroffe, C. D. (1985) (p. 14)
Voucher cited: Woodroffe 192 (CHR) |
|
|
United States (other Pacific offshore islands)
Midway Atoll |
Midway Atoll |
introduced
invasive cultivated abo. intro. |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1873) |
|
United States (other Pacific offshore islands)
Wake Islands |
Wake Island |
introduced
cultivated |
Fosberg, F. R. (1959) (p. 17)
Voucher cited: Fosberg 34449 Persisting but rare in 1952. |
|
United States (other Pacific offshore islands)
Wake Islands |
Wake Island |
introduced
|
Fosberg, F. R./Sachet, M.-H. (1969) (p. 13)
Not seen in 1961 or 1963. |
|
United States (other Pacific offshore islands)
Wake Islands |
Wake Island |
introduced
invasive cultivated |
Fosberg, F. R. (1955) (p. 17)
Voucher cited: Fosberg 34449 (US) Persisting from cultivation. |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Honduras
Honduras |
Honduras (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 193) | |
|
Indonesia
Indonesia |
Indonesia (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 193) | |
|
Taiwan
Taiwan |
Taiwan Island | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 193) | |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia (Indian Ocean offshore islands)
Christmas Island Group |
Christmas Island |
introduced
cultivated |
Swarbrick, J. T. (1997) (p. 116)
Naturalized |
|
British Indian Ocean Territory
Chagos Archipelago |
Diego Garcia Island |
Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 156)
"Recorded by Bourne (Hemsley 1884, p. 334)". |
|
Comments: Ipomoea batatas is a early introduction throughout the Pacific. The above lists documented locations, but it is undoubtedly present on many more islands.
Introduced and naturalized in undisturbed habitats in the Galápagos Islands.
Additional information: Additional online information about Ipomoea batatas is available from the Hawaiian Ecosystems at Risk project (HEAR).
Taxonomic information about Ipomoea batatas may be available from the Germplasm Resources Information Network (GRIN).
References:
Anderson, Donald. 1951. The plants of Arno Atoll, Marshall Islands. Atoll Research Bulletin No. 7. Pacific Science Board, National Academy of Sciences, Washington. 4 pp. + 7 pp. appendix.
Charles Darwin Research Station. 2005. CDRS Herbarium records.
Doty, Maxwell S. 1954. Floristic and Ecological Notes on Raroia Atoll, Tuamotus. Land biota: vascular plants. Atoll Research Bulletin No. 33. Pacific Science Board, National Academy of Sciences, Washington. 41 pp.
Ellison, Joanna C. 1990. Vegetation and floristics of the Tongatapu outliers. Atoll Research Bulletin No. 332. Smithsonian Institution, Washington. 36 pp.
Fosberg, F. R. 1955. Northern Marshall Islands Expedition, 1951-1952. Land biota: vascular plants. Atoll Research Bulletin No. 39. Pacific Science Board, National Academy of Sciences, Washington. 22 pp.
Fosberg, F. R. 1959. Vegetation and flora of Wake Island. Atoll Research Bulletin 67. Pacific Science Board, National Academy of Sciences, Washington. 20 pp.
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