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(Retz.) C. E. Hubb., Euphorbiaceae |
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Present on Pacific Islands? yes
Primarily a threat at high elevations? no
Common name(s): [more details]
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Chinese: ze qi |
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English: sun spurge, wolf's-milk |
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French: euphorbe reveil matin, tithymale |
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Japanese: todaigusa |
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Spanish: lecherina, lecheruela, lechetrezna comun, mamona, pichoa, pichoga |
Habit: herb
Description: "An annual monoecious herb with milky sap throughout; branched taproot up to 80 cm long; stems simple or branched at base, yellowish-green or sometimes purplish, glabrous, erect or ascending 15 to 60 cm tall, bearing terminal flower clusters; leaves alternate, 1 to 3 cm long, 1 to 1.5 cm wide, terminal leaves often in whorls of 5, simple, sessile, serrulate, obovate or rounded at apex, margins entire or finely toothed at apex; inflorescence composed of 5 umbel-like cyathia (cup-like receptacles, each containing a flowering group), subtended by a whorl of 5 leaves, smaller than stem leaves and not as narrow at base; each branch divides into 3 branches of second order and then 2 of third order with terminal flower cluster; involucre urn-shaped, with 4 rounded glands; flowers without petals, unisexual, flowering group with a single, central pistillate flower with a gynoecium: a long stalk with 3 styles and 3-celled ovary that extends beyond involucre after anthesis; surrounded by 10 to 12 simple staminate flowers each consisting of single stamen with small bract at base; fruit a dry, dehiscent schizocarp 4 to 5 mm in diameter with 3 carpels; at maturity carpels open violently; seed obovate to ovate in outline, circular in cross-section, 2 mm long, 1.5 mm wide, dark brown to orange-brown, with surface roughened by raised fine-meshed network; hilum kidney shaped, white and very conspicuous. This species is recognized by its milky sap throughout, flowers in terminal umbels of branched cymes, each surrounded by a whorl of five leaves, and the sharply reticulate seed surface" (Holm et al., 1997; p. 353).
"Annual, stems ascending, 0.5-6 dm high; leaves all obovate and very rounded or retuse at the end, finely serrate, smooth or a little hairy, those of the stem cuncate; umbel divided into usually 5 4 rays, then into 3, or at length simply forked; floral leaves narrowed to base; glands orbicular or elliptical, yellow, stalked; capsule smooth and even; seeds with coarse honeycomb-like reticulations" (Fernald, 1970; pp. 967-968).
Habitat/ecology: Waste places and dry open soil (Fernald, 1970; pp. 967-968). In New Caledonia, "mauvaise herbe peu commune des jardins" (MacKee, 1994; p. 51). "In Chile this species grows in the following environmental conditions: Medium altitude up to the timber line; low altitude, interior valleys; coastal mountains, 500-2000 m; coastal areas, 0-500 m. The plant grows in water or it has its roots within a permanent water course, this corresponds to marshes, bogs, water courses, lake and river shores; humid areas, with almost constant rainfall, short dry periods are possible (generally not longer than 1 month). Fully exposed to the sun, level areas or slopes facing north; some shadow, some protection against direct sunlight, some shadow from vegetation, filtering about 20-40% of light" (Chileflora).
Propagation: Seed
Native range: Europe, temperate and tropical Asia, northern Africa; naturalized elsewhere (GRIN).
Presence:
| Pacific | |||
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Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
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Chile (offshore islands)
Juan Fernández Islands |
Isla Más a Tierra (Robinson Crusoe Island) |
introduced
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Cuevas, Jaime G./Marticorena, Alicia/Cavieres, Lohengrin A. (2004) (p. 535)
Voucher cited: CONC 158065, JF 186 |
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Chile (offshore islands)
Juan Fernández Islands |
Isla Más a Tierra (Robinson Crusoe Island) |
introduced
invasive cultivated |
Atkinson, Rachel/Sawyer, John (2011) |
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Chile (offshore islands)
Juan Fernández Islands |
Isla Más a Tierra (Robinson Crusoe Island) |
introduced
invasive cultivated |
Danton, Philippe/Perrier, Christophe/Martinez Reyes, Guido (2006) (p. 553) |
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New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive |
Tassin, Jacques (2005) |
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New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive |
MacKee, H. S. (1994) (p. 51)
Voucher cited: MacKee 40818 |
| Pacific Rim | |||
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Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
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Australia
Australia (continental) |
New South Wales |
introduced
invasive |
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 351) |
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Australia
Australia (continental) |
Queensland |
introduced
invasive |
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 351) |
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China
China |
China (People's Republic of) |
native
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Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2013)
Fields, roadsides, scrub, margins of mixed forests. Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Liaoning, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan, Xinjiang, Yunnan, Zhejiang. |
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China
China |
Hong Kong |
introduced
cultivated |
Wu, Te-lin (2001) (p. 180) |
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Japan
Japan |
Japan |
native
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U.S. Dept. Agr., Agr. Res. Serv. (2013) |
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North Korea
North Korea |
North Korea (Democratic People's Republic of) |
native
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U.S. Dept. Agr., Agr. Res. Serv. (2013) |
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South Korea
South Korea |
South Korea (Republic of) |
native
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U.S. Dept. Agr., Agr. Res. Serv. (2013) |
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New Zealand
New Zealand |
New Zealand (country) |
introduced
invasive |
Webb, C. J./Sykes, W. R./Garnock-Jones, P. J. (1988) (p. 628)
"Common in gardens, waste places and crops, occasional in riverbeds". |
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South America (Pacific rim)
South America (Pacific rim) |
Chile (Republic of) |
introduced
invasive |
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 351) |
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Taiwan
Taiwan Island |
Taiwan Island |
native
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U.S. Dept. Agr., Agr. Res. Serv. (2013) |
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Vietnam
Vietnam |
Vietnam (Socialist Republic of) |
native
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U.S. Dept. Agr., Agr. Res. Serv. (2013) |
Control: If you know of control methods for Euphorbia helioscopia, please let us know.