|
L., Verbenaceae |
|
Present on Pacific Islands? yes
Primarily a threat at high elevations? no
Risk assessment results: High risk, score: 13 (Go to the risk assessment)
Other Latin names: Duranta repens L.
Common name(s): [more details]
|
Chinese: jia lian qiao |
|
English: duranta, golden dewdrop, golden eardrops, pigeon-berry, sky-flower |
|
French: durante dressée |
|
Marshallese: jab meloklok |
|
Spanish: cuentas de oro, San Jacinto |
|
Tongan: māvaetangi, olive |
Habit: shrub
Description: "Shrub to 4-6 m tall; spines axillary or suppressed; leaves opposite or 3 in a whorl, elliptic or ovate, rarely elliptic-obovate, 1.5-7 cm long, entire or distally serrate, acute, very short-petiolate; racemes 5-15 cm long; flowers subsessile; calyx 3-4 mm long; corolla pale purplish, with 2 slightly small lobes each with a purple stripe, and 3 slightly larger plain lobes; fruit globose, 6-12 mm broad, yellow, beaked" (Stone, 1970; p. 389).
Habitat/ecology: "Golden dewdrop grows wild mostly in dry coastal areas (750 to 900 mm of annual precipitation in Puerto Rico) from near sea level to over 100 m in elevation (Little and others 1974). Because it is moderately intolerant of shade and does not compete well with taller vegetation, golden dewdrop is usually found in rocky or sandy areas with low shrubs and sparse grass and herbs. Although it is more common on limestone, the shrub also grows in areas with igneous rocks. The species tolerates light to moderate salt spray. In natural stands, golden dewdrop may be found in small tangled stands or as occasional plants. Plants under droughty and infertile natural conditions tend to be short (about 1 m in height). Where cultivated in deep, moist, fertile soils, golden dewdrop grows faster and to larger sizes. As an ornamental, it can be grown in moderate shade at elevations up to 1,300 m (Bruggeman 1974)" (Wildland shrubs of the United States and its territories). Dry to moist disturbed areas. Common along roadsides and in thickets, 450-5500 feet in Jamaica (Adams, 1972; pp. 632-633).
Propagation: Seed
Native range: Tropical America, widely cultivated and naturalized in the tropics.
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
American Samoa
Manua Islands |
Tau Island |
introduced
cultivated |
Space, James C./Flynn, Tim (2000) (p. 45) |
|
American Samoa
Tutuila Islands |
Tutuila Island |
introduced
cultivated |
Space, James C./Flynn, Tim (2000) (p. 35) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235) |
|
Cook Islands
Southern Cook Islands |
Rarotonga Island |
cultivated
|
Sykes, Bill (year unknown) |
|
Ecuador (Galápagos Islands)
San Cristóbal Group |
San Cristóbal Island |
introduced
cultivated |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Santa Cruz Group |
Santa Cruz Island |
introduced
cultivated |
Charles Darwin Foundation (2008) |
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. (2000) (p. 27) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
cultivated |
Lorence, David H./Flynn, Timothy (2010) (p. 18)
Golden leaf cultivar. |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235) |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
|
Glassman, S. F. (1952) (p. 103)
Voucher cited: Takamatsu 628 (BISH) |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
cultivated |
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 100)
Voucher cited: M. Takamatsu 628 (BISH) |
|
Fiji
Fiji Islands |
Vanua Levu Island |
introduced
invasive cultivated |
Smith, Albert C. (1991) (pp. 176-177)
Voucher cited: DA 16959 Occasional plantation weed. |
|
Fiji
Fiji Islands |
Vanua Mbalavu Island |
introduced
invasive cultivated |
Smith, Albert C. (1991) (pp. 176-177)
Vouchers cited: DA L.13847, DA L.14390 Occasional plantation weed. |
|
Fiji
Fiji Islands |
Viti Levu Island |
introduced
invasive cultivated |
Smith, Albert C. (1991) (pp. 176-177)
Vouchers cited: Smith 5024, DA 6037, DA 12345, DA 16097 Occasional plantation weed. |
|
French Polynesia
Gambier Islands |
Mangareva (Gambier) Island |
introduced
cultivated |
St. John, Harold (1988) (p. 22)
Voucher cited: Fosberg 11089 (BISH) |
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
introduced
cultivated |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
introduced
cultivated |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 294)
Voucher cited: Florence 5027 |
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 5027 (PAP) Naturalisée |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
invasive cultivated |
Welsh, S. L. (1998) (p. 294)
Voucher cited: Moore 365, BRY 25393 |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 3484 (PAP), F.R. Fosberg & M.-H. Sachet 63209 (PAP) Naturalisée |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive cultivated |
Welsh, S. L. (1998) (p. 294)
Vouchers cited: Florence 2610, Grant 3847 |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive cultivated |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 2610 (PAP) Naturalisé. Ornementale commune à Tahiti. |
|
Guam
Guam Island |
Guam Island |
introduced
cultivated |
Stone, Benjamin C. (1970) (p. 505) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive |
Lorence, David H./Flynn, Timothy W./Wagner, Warren L. (1995) (p. 55)
Vouchers cited: D. Lorence 6624 (BISH, PTBG), T. Flynn & D. Harter 2018 (PTBG) Naturalized in secondary vegetation. |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive |
Herbarium Pacificum Staff (1998) (pp. 13-14)
Voucher cited: K. Thorne & H. Black 7862 (BISH, BRY) |
|
Marshall Islands
Ralik Chain |
Kwajalein (Kuwajleen) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235) |
|
Marshall Islands
Ralik Chain |
Kwajalein (Kuwajleen) Atoll |
introduced
cultivated |
Whistler, W. A./Steele, O. (1999) (p. 103) |
|
Marshall Islands
Ratak Chain |
Arno Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 235) |
|
Marshall Islands
Ratak Chain |
Arno Atoll |
Anderson, Donald (1951) (p. ii)
Voucher cited: Anderson 3760 |
|
|
Nauru
Nauru Island |
Nauru Island |
introduced
cultivated |
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 205) |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Lifou |
introduced
|
MacKee, H. S. (1994) (p. 139)
Vouhcer cited: Sarasin 791 (Z) |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
cultivated |
MacKee, H. S. (1994) (p. 139)
Vouchers cited: de Pompéry s.n., MacKee 24976 |
|
Palau
Palau (main island group) |
Angaur Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 103) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 101) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
cultivated |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 19) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 97) |
|
Palau
Palau (main island group) |
Malakal Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 97) |
|
Palau
Palau (main island group) |
Ngercheu Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 108)
Carp Resort |
|
Palau
Palau (main island group) |
Ngerkebesang Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 97) |
|
Palau
Palau (main island group) |
Peleliu Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 106) |
|
Palau
Sonsorol Group |
Sonsorol Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (pp. 15, 109) |
|
Philippines
Philippine Islands |
Philippine Islands |
introduced
cultivated |
Merrill, Elmer D. (1923) (p. 381)
Cultivated for ornamental purposes. |
|
Samoa
Western Samoa Islands |
Savaii Island |
introduced
cultivated |
Space, James C./Flynn, Tim (2002) (p. 70)
Voucher: Flynn 6961 (PTBG, US, SAMOA) |
|
Samoa
Western Samoa Islands |
Savaii Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 270)
Taxon name on voucher: Duranta erecta |
|
|
Samoa
Western Samoa Islands |
Upolu Island |
introduced
cultivated |
Space, James C./Flynn, Tim (2002) (p. 66) |
|
Tonga
Haapai Group |
Lifuka and Foa Islands |
introduced
cultivated |
Space, James C./Flynn, Tim (2001) (p. 72) |
|
Tonga
Tongatapu Group |
Eua Island |
introduced
cultivated |
Space, James C./Flynn, Tim (2001) (p. 60) |
|
Tonga
Tongatapu Group |
Eua Island |
introduced
invasive cultivated |
Yuncker, T. G. (1959) (p. 231)
Voucher cited: Yuncker 15499 An escape collected. |
|
Tonga
Tongatapu Group |
Tongatapu Island |
introduced
cultivated |
Space, James C./Flynn, Tim (2001) (p. 52) |
|
Vanuatu
New Hebrides Islands |
Vanuatu (Republic of) | Swarbrick, John T. (1997) (p. 91) | |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis (Uvea) Island |
introduced
cultivated |
Meyer, Jean-Yves (2007) (pp. 13, 38) |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia
Australia (continental) |
Queensland |
introduced
invasive |
Queensland Herbarium (2002) (p. 3) |
|
China
China |
China (People's Republic of) |
introduced
invasive cultivated |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
Usually cultivated or sometimes naturalized near farm houses; 200-400 m. Naturalized in Fujian, Guangdong, Guangxi, Hainan, Hunan, Jiangxi, Zhejiang. |
|
China
China |
Hong Kong |
introduced
cultivated |
Wu, Te-lin (2001) (p. 230)
Hedges. |
|
Colombia
Colombia |
Colombia (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Costa Rica
Costa Rica |
Costa Rica (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Ecuador (Mainland)
Ecuador |
Ecuador (Republic of) (continental) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
El Salvador
El Salvador |
El Salvador (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Guatemala
Guatemala |
Guatemala (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Honduras
Honduras |
Honduras (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Mexico
Mexico |
Mexico (United Mexican States) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Nicaragua
Nicaragua |
Nicaragua (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Panama
Panama |
Panama (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Perú
Perú |
Perú (Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Singapore
Singapore |
Singapore (Republic of) |
introduced
cultivated |
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 37)
Cultivated only |
|
Taiwan
Taiwan |
Taiwan Island |
introduced
invasive cultivated |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
Usually cultivated or sometimes naturalized near farm houses; 200-400 m. |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia (Indian Ocean offshore islands)
Christmas Island Group |
Christmas Island |
introduced
cultivated |
Swarbrick, J. T. (1997) (p. 112) |
|
La Réunion (France)
La Réunion Island |
La Réunion Island |
introduced
cultivated |
Comité français de l'Union Internationale pour la Conservation de la Nature en France (2010)
Naturalisée, non envahissant. |
|
French Territory of Mayotte
Mayotte Islands |
Mayotte Island |
introduced
invasive cultivated |
Comité français de l'Union Internationale pour la Conservation de la Nature en France (2010) |
Comments: Naturalizing in Hawaii.
Additional information:
Fact sheet from "Wildland
shrubs of the United States and its territories: thamnic descriptions" (PDF format).
Additional online information about Duranta erecta is available from the Hawaiian Ecosystems at Risk project (HEAR).
Information about Duranta erecta as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).
Taxonomic information about Duranta erecta may be available from the Germplasm Resources Information Network (GRIN).
References:
Adams, C. D. 1972. Flowering plants of Jamaica. University of the West Indies, Mona, Jamaica. 848 pp.
Anderson, Donald. 1951. The plants of Arno Atoll, Marshall Islands. Atoll Research Bulletin No. 7. Pacific Science Board, National Academy of Sciences, Washington. 4 pp. + 7 pp. appendix.
Charles Darwin Foundation. 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Charles Darwin Foundation, Galapagos, Ecuador.
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species. Raffles Museum of Biodiversity Research, National University of Singapore. 273 pp.
Comité français de l'Union Internationale pour la Conservation de la Nature en France. 2010. Les espéces envahissantes en outre-mer (online resource).
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. 2011. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP). (online resource).
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.
Francis, John K., ed. 2009. Wildland Shrubs of the United States and its Territories: Thamnic Descriptions General Technical Report IITF-WB-1 U.S. Department of Agriculture, Forest Service International Institute of Tropical Forestry and Shrub Sciences Laboratory (online resource).
Glassman, S. F. 1952. The flora of Ponape. Bishop Mus. Bull. 209: 1-152.
Herbarium Pacificum Staff. 1998. New Hawaiian plant records for 1997. In: Evenhuis, Neal L. and Miller, Scott, E., eds. Records of the Hawaii Biological Survey for 1997. Part 2: Notes. Bishop Museum Occasional Papers. 56:8-14.
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. 2010. Checklist of the vascular plants of Pohnpei with local names and uses. Allertonia, in press. National Tropical Botanical Garden, Lawai, Hawaii. 146 pp.
Humphries, S. E./Stanton, J. P. 1992. Weed assessment in the wet tropics world heritage area of north Queensland. Report to The Wet Tropics Management Agency. 75 pp. plus plates.
Lorence, David H./Flynn, Timothy. 2010. Checklist of the plants of Kosrae. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawaii. 26 pp.
Lorence, David H./Flynn, Timothy W./Wagner, Warren L. 1995. Contributions to the flora of Hawaii. III. New additions, range extensions, and rediscoveries of flowering plants. In: Evenhuis, Neal L. and Miller, Scott, E., eds. Records of the Hawaii Biological Survey for 1994. Part 1: Articles. Bishop Museum Occasional Papers. 41:19-58.
Lorence, David H./Wagner, Warren L. 2008. Flora of the Marquesas Islands. National Tropical Botanical Garden and the Smithsonian Institution. Online database.
MacKee, H. S. 1994. Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie. Muséum National d'Histoire Naturelle, Paris, 164 p.
Merrill, Elmer D. 1923. An enumeration of Philippine flowering plants, vol. 3 [reprint]. Bureau of Printing, Manila. 628 pp.
Meyer, Jean-Yves. 2007. Rapport de mission sur l'Ile d'Uvea (Wallis & Futuna) du 6 au 17 Novembre 2007: Inventaire preliminaire de la flore vasculaire secondaire [unpublished report]. Délégation à la Recherche, Ministère de l'Education, l'Enseignement Supérieur et la Recherche, B.P. 20981 Papeete, Tahiti, Polynésie française. 39 pp.
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.). 2002. Voucher specimen #PTBG270(Tim Flynn 6961).
Queensland Herbarium. 2002. Invasive Naturalised Plants in Southeast Queensland, alphabetical by genus. Modified from: Batianoff, George N. and Butler, Don W. (2002). Assessment of Invasive naturalized plants in south-east Queensland. Appendix. Plant Protection Quarterly 17, 27-34. 11 pp.
Smith, Albert C. 1991. Flora Vitiensis nova: a new flora of Fiji. National Tropical Botanical Garden, Lawai, Kauai, Hawaii. Volume 5. 626 pp.
Space, James C./Flynn, Tim. 2000. Observations on invasive plant species in American Samoa. USDA Forest Service, Honolulu. 51 pp.
Space, James C./Flynn, Tim. 2001. Report to the Kingdom of Tonga on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 78 pp.
Space, James C./Flynn, Tim. 2002. Report to the Government of Samoa on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 83 pp.
Space, James C./Lorence, David H./LaRosa, Anne Marie. 2009. Report to the Republic of Palau: 2008 update on Invasive Plant Species. USDA Forest Service, Hilo, Hawaii. 227 pp.
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Mazawa, Thomas R. 2000. Invasive plant species in Chuuk, Federated States of Micronesia. USDA Forest Service, Honolulu. 41 pp.
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar. 2003. Report to the Republic of Palau on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 174 pp.
St. John, Harold. 1988. Census of the Flora of the Gambier Islands, Polynesia. Pacific Plant Studies 43. Honolulu, Hawaii. 34 pp. [Privately published].
Stone, Benjamin C. 1970. The flora of Guam. Micronesica 6:1-659.
Swarbrick, J. T. 1997. Environmental weeds and exotic plants on Christmas Island, Indian Ocean: a report to Parks Australia. 101 pp. plus appendix.
Swarbrick, John T. 1997. Weeds of the Pacific Islands. Technical paper no. 209. South Pacific Commission, Noumea, New Caledonia. 124 pp.
Sykes, Bill. 0. Bill Sykes, pers. com.
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. 1994. The flora of Nauru. Smithsonian Institution, Washington. Atoll Research Bulletin 392:1-223.
U.S. Dept. Agr., Agr. Res. Serv. 2011. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawaii Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).
Welsh, S. L. 1998. Flora Societensis: A summary revision of the flowering plants of the Society Islands. E.P.S. Inc., Orem, Utah. 420 pp.
Whistler, W. A. 1991. The ethnobotany of Tonga: The plants, their Tongan names, and their uses. Bishop Museum Bulletin in Botany 2. Bishop Museum Press, Honolulu. 155 pp. ISBN 0-930897-57-9.
Whistler, W. A./Steele, O. 1999. Botanical survey of the United States of America Kwajalein Atoll (USAKA) Islands. Prepared for Oak Ridge Institute for Science and Education and the U. S. Army Environmental Center. 111 pp.
Wiggins, I. L./Porter, D. M. 1971. Flora of the Galapágos Islands. Stanford University Press. 998 pp.
Wu, Te-lin. 2001. Check List of Hong Kong Plants. Hong Kong Herbarium and the South China Institute of Botany. Agriculture, Fisheries and Conservation Department Bulletin 1 (revised). 384 pp.
Yuncker, T. G. 1959. Plants of Tonga. Bishop Museum Bull. 220. Bishop Museum Press, Honolulu. 343 pp.
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong. 2011. Flora of China (online resource).