Pacific Island Ecosystems at Risk (PIER)

  [   PIER species lists  ]   [   PIER home  ]

Desmodium heterocarpon
(L.) DC., Fabaceae
Click on an image for links to BIGGER PICTURES


Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Other Latin names:  Desmodium ovalifolium (Prain) Wall. ex Merr.; Desmodium polycarpon (Poir.) DC.; Meibomia heterocarpos (L.) Kuntze

Common name(s): [more details]

Chinese: jia di dou

English: Asian ticktrefoil, carpon desmodium

Mangarevan: piri piri, pua marino

Palauan: okula beluu

Pohnpeian: kaamalimal, kamelimel, kehmelmel en nansapw

Tahitian: ea ea, piripiri ‘aratita, piripiro, tia pipi, toharea

Tongan: ngingie ‘uta

Habit:  herb

Description:  "Suberect subshrubs usually 5-15 dm tall; stems usually many-branched, appressed pubescent in upper part.  Leaves (1-) 3-foliolate, leaflets subcoriaceous, usually elliptic, terminal one 2.5-6 (-8) cm long, (1-) 1.5-2.5 (-3) cm wide, upper surface subglabrous, lower surface sericeous with appressed hairs, apex obtuse, petioles (1-) 1.5-3.5 cm long.  Flowers numerous in racemose inflorescences 5-10 cm long, rachis densely pubescent with appressed, straight, yellowish hairs, pedicels (2-) 3-5 (-8) mm long; corolla pink to purplish red.  Pods narrowly oblong, 4-8-jointed, 1-2.8 cm long, densely hooked pubescent, articles quadrate, 2.5-3 mm long, 2-3 mm wide."

Desmodium heterocarpon (L.) DC. subsp. heterocarpon:  inflorescence rachis densely pubescent with spreading hooked hairs.

Desmodium heterocarpon var. strigosum: inflorescence rachis densely pubescent with appressed, straight, non-hooked hairs.  (Wagner et al., 1999; p. 667).

Habitat/ecology:  In Fiji, "occurring from near sea level to about 200 m in pastures and secondary growth, along roadsides, on waste land, and sometimes thoroughly naturalized in dry forest" (Smith, 1985; pp. 192-193).  "Uncommon to occasional in sunny, disturbed places" (Whistler, 1988; p. 23).

Propagation:  Seed

Native range:  Old World (Wagner et al., 1999; p. 667).  "Variety strigosum appears indigenous from southeastern Asia throughout Malesia to Micronesia and possibly to New Caledonia; eastward in the Pacific as far as the Tuamotus and Hawai'i. It was presumably an inadvertent aboriginal introduction." (Smith, 1985; pp. 192-193)  "An aboriginal introduction to Polynesia" (Whistler, 1988; p. 23).

Presence:

Pacific
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
American Samoa
Manu‘a Islands
Ofu Island   Whistler, W. A. (1990)
Vouchers cited: Garber 979, Yuncker 9469, Whistler 2995
American Samoa
Manu‘a Islands
Ofu Island   Yuncker, T. G. (1945) (p. 41)
Vouchers cited: Yuncker 9469 (BISH), Garber 979 (BISH)
American Samoa
Manu‘a Islands
Olosega Island   Whistler, W. A. (1990)
Voucher cited: Whistler 3060
American Samoa
Manu‘a Islands
Olosega Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2001) (voucher ID: PTBG 35046)
Taxon name on voucher: Desmodium heterocarpum (L.) DC.
American Samoa
Manu‘a Islands
Olosega Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1991) (voucher ID: PTBG 10498)
Taxon name on voucher: Desmodium heterocarpon
American Samoa
Manu‘a Islands
Ta‘u Island   Whistler, W. A. (1990)
Vouchers cited: Garber 603, Garber 634, Garber 645a, Yuncker 9083, Whistler 1391
American Samoa
Manu‘a Islands
Ta‘u Island   Yuncker, T. G. (1945) (p. 41)
Vouchers cited: Yuncker 9083 (BISH), Garber 603 (BISH), Garber 634 (BISH), Garber 645a (BISH)
American Samoa
Manu‘a Islands
Ta‘u Island introduced
Ragone, Diane/Lorence, David H. (2003) (p. 49)
American Samoa
Tutuila Islands
Tutuila Island   Swarbrick, John T. (1997) (p. 91)
American Samoa
Tutuila Islands
Tutuila Island introduced
abo. intro.
Whistler, W. A. (1980) (p. 131)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Northern Mariana Islands   Swarbrick, John T. (1997) (p. 91)
Cook Islands
Southern Cook Islands
Mangaia Island introduced
invasive
Space, James C./Flynn, Tim (2002) (p. 16)
Cook Islands
Southern Cook Islands
Ma‘uke Island   McCormack, Gerald (2013)
?
Cook Islands
Southern Cook Islands
Ma‘uke Island introduced
invasive
Space, James C./Flynn, Tim (2002) (p. 16)
Voucher: Flynn 7086 (PTBG, BISH, NY, US)
Cook Islands
Southern Cook Islands
Ma‘uke Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 385)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Federated States of Micronesia
Chuuk Islands
Weno (Moen) Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109)
var. strigosum f. substrigosum
Federated States of Micronesia
Kosrae Island
Kosrae Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2005) (voucher ID: PTBG 42612)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109)
var. strigosum f. substrigosum
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   Glassman, S. F. (1952) (p. 76)
Vouchers cited: Glassman 2316 (US), Glassman 2445 (US), Kanehira 1574 (NY)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island native
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 92)
Vouchers cited: M.V.C. Falanruw MVCF3223 (BISH), M.V.C. Falanruw MVCF5405 (BISH), R. Gallen 38 (NY, PTBG), S.F. Glassman 2316 (BISH), S.F. Glassman 2445 (BISH), E.Y. Hosokawa 3502 (BISH), T. Hosokawa 5830 (BISH), T. Hosokawa 9569 (BISH), R. Kanehira 1574 (BISH), S. Perlman 20295 (PTBG), P. Raynor 158 (NY, PTBG), M. Takamatsu 740 (BSIH)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1985) (voucher ID: PTBG 23357)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Federated States of Micronesia
Yap Islands
Yap (Waqab) Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109)
var. strigosum f. substrigosum
Fiji
Fiji Islands
Koro Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Voucher cited: DA 11803
Fiji
Fiji Islands
Matuku Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Voucher cited: Bryan 260
Fiji
Fiji Islands
Mbengga Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Voucher cited: DA 9619
Fiji
Fiji Islands
Ngau Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Voucher cited: Milne 150
Fiji
Fiji Islands
Ovalau Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Voucher cited: Seemann 111
Fiji
Fiji Islands
Taveuni Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Vouchers cited: DA 2564, Gillespie 4774
Fiji
Fiji Islands
Vanua Levu Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Vouchers cited: Smith 6811, DA 11779, DA 13137
Fiji
Fiji Islands
Viti Levu Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Vouchers cited: Greenwood 452, DA 513, DA 10005, B. & H. Parham 35, DA 1052
Fiji
Fiji Islands
Wakaya Island introduced
invasive
Smith, Albert C. (1985) (pp. 192-193)
var. strigosum. Voucher cited: Milne 377
French Polynesia
Gambier Islands
Akamaru Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Gambier Islands
Akamaru Island native
St. John, Harold (1988) (p. 15)
Voucher cited: St. John 14698 (BISH)
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen. Voucher cited: B. Huguenin s.n. (PAP)
Adventice
French Polynesia
Gambier Islands
Mangareva (Gambier) Island native
St. John, Harold (1988) (p. 15)
Vouchers cited: Agassiz 105 (GH), Fosberg 11065 (BISH), Fosberg 11098 (BISH), Fosberg 11117 (BISH), St. John 14482 (BISH)
French Polynesia
Gambier Islands
Taravai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Gambier Islands
Taravai Island native
St. John, Harold (1988) (p. 15)
Voucher cited: St. John 14779 (BISH)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Marquesas Islands
Hatuta'a (Hatutu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
Var. strigosum Meeuwen
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
Var. strigosum Meeuwen
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 7288 (PAP)
Adventice
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
Var. strigosum Meeuwen
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Society Islands
Bora Bora Island introduced
Welsh, S. L. (1998) (p. 149)
Voucher cited: Grant 5026
French Polynesia
Society Islands
Bora Bora Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Society Islands
Huahine Island introduced
Welsh, S. L. (1998) (p. 149)
Voucher cited: D. Bare 9
French Polynesia
Society Islands
Huahine Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: D. Baré 8 (PAP)
Adventice
French Polynesia
Society Islands
Maupiti (Maurua) Island introduced
Fosberg, F. R./Sachet, M.-H. (1987) (p. 35)
Not seen in 1985. Voucher cited: Raynal 17860.
French Polynesia
Society Islands
Mehetia (Mahetia, Meetia) Island introduced
Welsh, S. L. (1998) (p. 149)
Voucher cited: Grant 4744
French Polynesia
Society Islands
Mehetia (Mahetia, Meetia) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Society Islands
Moorea Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
Welsh, S. L. (1998) (p. 149)
Vouchers cited: Moore 3, BRY 25296, BRY 26701
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen. Voucher cited: J. Florence 11857 (PAP)
Adventice
French Polynesia
Society Islands
Tahiti Island introduced
Welsh, S. L. (1998) (p. 149)
Voucher cited: Florence 8064
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen. Vouchers cited: J. Florence 8064 (PAP), J. Florence 9893 (PAP), J. Florence & S. Jourdan 10719 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Makatea (Ma‘atea) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Austral (Tubuai) Islands
Raivavae (Raevavae) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Austral (Tubuai) Islands
Rimatara (Rimitara) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
French Polynesia
Austral (Tubuai) Islands
Tubuai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. strigosum M.S.K. Meeuwen
Adventice
State of Hawaii
Hawaiian Islands
O‘ahu Island introduced
Wester, Lyndon (1992) (p. 138)
Japan (offshore islands)
Ogasawara (Bonin) Islands
Bonin (Ogasawara) Islands   Kato, Hidetoshi (2007)
Kiribati
Kiribati
Kiribati (Republic of) introduced
Fosberg, F. R./Sachet, M.-H. (1987) (p. 16)
Marshall Islands
Marshall Islands
Marshall Islands   Swarbrick, John T. (1997) (p. 91)
New Caledonia
New Caledonia
New Caledonia Islands native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Niue
Niue
Niue Island introduced
Sykes, W. R. (1970) (pp. 151-152)
Voucher cited: CHR 150536
Niue
Niue
Niue Island introduced
invasive
abo. intro.
Whistler, W. A. (1988) (p. 23)
Palau
Palau (Belau ) (main island group)
Babeldaob Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109)
var. strigosum f. strigosum.
Palau
Palau (Belau ) (main island group)
Babeldaob Island native
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. Voucher specimen)
var. strigosum. Voucher: BMW 6557 (BISH, BRI)
Palau
Palau (Belau ) (main island group)
Babeldaob Island native
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. Field notes and voucher.)
var. strigosum. Voucher: Lorence 9911 (PTBG, BNM, NY, US)
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1966) (voucher ID: BNM 455)
Taxon name on voucher: Desmodium heterocarpon var heterocarpon (Willd.) DC.
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1981) (voucher ID: BNM 3149)
Taxon name on voucher: Desmodium heterocarpon (Willd.) DC.
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (2007) (voucher ID: BNM 4401)
Taxon name on voucher: Desmodium heterocarpon (L.) DC. var. strigosum v. Meeuwen
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1991) (voucher ID: BNM 9569)
Taxon name on voucher: Desmodium heterocarpon var. strigosum V. Meeuwen.
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1981) (voucher ID: BNM 3098)
Taxon name on voucher: Desmodium heterocarpon var. strigosum v. Meeuwen
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1978) (voucher ID: BNM 1255)
Taxon name on voucher: Desmodium heterocarpon var. strigosum v. Meeuwen
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1982) (voucher ID: BNM 3195)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau
Palau (Belau ) (main island group)
Babeldaob Island   Beleu National Museum (Republic of Palau. Koror.) (1978) (voucher ID: BNM 10153)
Taxon name on voucher: Desmodium heterocarpon cf. var. strigosum Meeuw.
Palau
Palau (Belau ) (main island group)
Babeldaob Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 204)
Taxon name on voucher: Desmodium heterocarpon (L.) DC.
Palau
Palau (Belau ) (main island group)
Babeldaob Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1966) (voucher ID: PTBG 22484)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau
Palau (Belau ) (main island group)
Babeldaob Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1983) (voucher ID: PTBG 22783)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau
Palau (Belau ) (main island group)
Babeldaob Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1981) (voucher ID: PTBG 22784)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109)
var. strigosum f. strigosum.
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   Beleu National Museum (Republic of Palau. Koror.) (1967) (voucher ID: BNM 456)
Taxon name on voucher: Desmodium heterocarpon
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1965) (voucher ID: PTBG 22782)
Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau
Palau (Belau ) (main island group)
Ngerkebesang Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109)
var. strigosum f. strigosum.
Palau
Palau (Belau ) (main island group)
Peleliu (Beliliou) Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Papua New Guinea
Bismarck Archipelago
Bismarck Archipelago native
ILDIS Co-ordinating Centre (2013)
Papua New Guinea
Bismarck Archipelago
Bismarck Archipelago   Peekel, P. G. [translated by E. E. Henty] (1984) (p. 237)
Frequent, in grassy places, at roadsides and in scrub.
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Philippines
Philippine Islands
Philippine Islands native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Philippines
Philippine Islands
Philippine Islands   Merrill, Elmer D. (1923) (pp. 285-286)
In thickets, open grasslands, etc., at low and medium altitudes; common.
Solomon Islands
Solomon Islands
Solomon Islands native
ILDIS Co-ordinating Centre (2013)
Solomon Islands
Solomon Islands
Solomon Islands introduced
invasive
Hancock, I. R./Henderson, C. P. (1988) (p. 65)
Tonga
Ha‘apai Group
Kao Island probably introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 29)
Vouchers cited: Yuncker 15871, Buelow 741
Tonga
Ha‘apai Group
Kao Island   Yuncker, T. G. (1959) (p. 140)
Voucher cited: Yuncker 15871
Tonga
Ha‘apai Group
Tofua Island probably introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 29)
Vouchers cited: Hotta 4331, Scarth-Johnson 75, Buelow 599
Tonga
Tonga Islands
Tonga Islands   Yuncker, T. G. (1959) (p. 140)
"Frequent throughout Tonga"
Tonga
Tongatapu Group
‘Eua Island probably introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 29)
Vouchers cited: Lister s.n., Yuncker 15372, Sykes 655, Sykes 668, Whistler 7414
Tonga
Tongatapu Group
‘Eua Island   Yuncker, T. G. (1959) (p. 140)
Vouchers cited: Yuncker 15372, Yuncker 15654
Tonga
Tongatapu Group
Tongatapu Island   Ellison, Joanna C. (1990) (p. 29)
On outlier islet of Polo‘a.
Tonga
Vava‘u Group
Late Island probably introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 29)
Voucher cited: Buelow & sykes 212
Tonga
Vava‘u Group
Niuafo‘ou Island probably introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 29)
Vouchers cited: Yuncker 16012, Buelow 2209
Tonga
Vava‘u Group
Niuafo‘ou Island   Yuncker, T. G. (1959) (p. 140)
Voucher cited: Yuncker 16012
Tonga
Vava‘u Group
Tafahi Island probably introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 29)
Vouchers cited: Hurlimann 524, Whistler 6236
Tonga
Vava‘u Group
Vava‘u Island   Yuncker, T. G. (1959) (p. 140)
Cited: Crosby.
Tuvalu
Tuvalu (Ellis) Islands
Tuvalu Islands   Swarbrick, John T. (1997) (p. 91)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis (‘Uvea) Island introduced
invasive
Meyer, Jean-Yves (2007) (p. 34)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis and Futuna Islands   Whistler, W. A. (1988) (p. 23)
Pacific Rim
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Australia
Australia (continental)
Queensland native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Cambodia
Cambodia
Cambodia (Kingdom of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
China
China
China (People's Republic of) native
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2013)
"Grasslands, grassy slopes, watersides, thickets, forests; 300-1800 m".
China
China
Hong Kong native
Wu, Te-lin (2001) (p. 143)
Indonesia
Indonesia
Indonesia (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Indonesia
Indonesia
West Papua Province (Indonesia) (western New Guinea Island) (formerly Irian Jaya) native
ILDIS Co-ordinating Centre (2013)
Japan
Japan
Japan (country) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Honshu, Kyushu, Shikoku
Japan
Ryukyu Islands
Ryukyu Islands native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Malaysia
Malaysia
Malaysia (country of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Singapore
Singapore
Singapore (Republic of) uncertain if introduced
invasive
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 33)
Weed of uncertain origin
South America (Pacific rim)
South America (Pacific rim)
Colombia (Republic of) introduced
ILDIS Co-ordinating Centre (2013)
South America (Pacific rim)
South America (Pacific rim)
Perú (Republic of) native
ILDIS Co-ordinating Centre (2013)
Taiwan
Taiwan Island
Taiwan Island native
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2013)
"Grasslands, grassy slopes, watersides, thickets, forests; 300-1800 m".
Thailand
Thailand
Thailand (Kingdom of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Vietnam
Vietnam
Vietnam (Socialist Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Indian Ocean
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Seychelles
Seychelles Islands
Seychelles Islands native
ILDIS Co-ordinating Centre (2013)

Comments:  Cook Islands subsp. heterocarpon, all other locations var. strigosum.

Previously collected in Hawai‘i but present status unknown (Wagner et al., 1999; p. 667).


Need more info? Have questions? Comments? Information to contribute? Contact PIER! (pier@hear.org)

  [   PIER species lists  ]   [   PIER home  ]

This page was created on 3 AUG 2002 and was last updated on 15 OCT 2006.