Desmodium heterocarpon (L.) DC., Fabaceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Common name(s): [more details]

Chinese: jia di dou

English: Asian ticktrefoil

Mangarevan: piri piri, pua marino

Palauan: okula beluu

Pohnpeian: kaamalimal, kamelimel, kehmelmel en nansapw

Tahitian: ea ea, piripiri ‘aratita, piripiro, tia pipi, toharea

Tongan: ngingie ‘uta

Habit:  herb

Description:  "Suberect subshrubs usually 5-15 dm tall; stems usually many-branched, appressed pubescent in upper part.  Leaves (1-) 3-foliolate, leaflets subcoriaceous, usually elliptic, terminal one 2.5-6 (-8) cm long, (1-) 1.5-2.5 (-3) cm wide, upper surface subglabrous, lower surface sericeous with appressed hairs, apex obtuse, petioles (1-) 1.5-3.5 cm long.  Flowers numerous in racemose inflorescences 5-10 cm long, rachis densely pubescent with appressed, straight, yellowish hairs, pedicels (2-) 3-5 (-8) mm long; corolla pink to purplish red.  Pods narrowly oblong, 4-8-jointed, 1-2.8 cm long, densely hooked pubescent, articles quadrate, 2.5-3 mm long, 2-3 mm wide."

Desmodium heterocarpon (L.) DC. subsp. heterocarpon:  inflorescence rachis densely pubescent with spreading hooked hairs.

Desmodium heterocarpon var. strigosum: inflorescence rachis densely pubescent with appressed, straight, non-hooked hairs.  (Wagner et al., 1999; p. 667).

Habitat/ecology:  In Fiji, "occurring from near sea level to about 200 m in pastures and secondary growth, along roadsides, on waste land, and sometimes thoroughly naturalized in dry forest" (Smith, 1985; pp. 192-193).  "Uncommon to occasional in sunny, disturbed places" (Whistler, 1988; p. 23).

Propagation:  Seed

Native range:  Old World (Wagner et al., 1999; p. 667).  "Variety strigosum appears indigenous from southeastern Asia throughout Malesia to Micronesia and possibly to New Caledonia; eastward in the Pacific as far as the Tuamotus and Hawai'i. It was presumably an inadvertent aboriginal introduction." (Smith, 1985; pp. 192-193)  "An aboriginal introduction to Polynesia" (Whistler, 1988; p. 23).

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
American Samoa Manu‘a Islands	Ofu Island		Whistler, W. A. (1990) Vouchers cited: Garber 979, Yuncker 9469, Whistler 2995
American Samoa Manu‘a Islands	Ofu Island		Yuncker, T. G. (1945) (p. 41) Vouchers cited: Yuncker 9469 (BISH), Garber 979 (BISH)
American Samoa Manu‘a Islands	Olosega Island		Whistler, W. A. (1990) Voucher cited: Whistler 3060
American Samoa Manu‘a Islands	Olosega Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2001) (voucher ID: PTBG 35046) Taxon name on voucher: Desmodium heterocarpum (L.) DC.
American Samoa Manu‘a Islands	Olosega Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1991) (voucher ID: PTBG 10498) Taxon name on voucher: Desmodium heterocarpon
American Samoa Manu‘a Islands	Ta‘u Island		Whistler, W. A. (1990) Vouchers cited: Garber 603, Garber 634, Garber 645a, Yuncker 9083, Whistler 1391
American Samoa Manu‘a Islands	Ta‘u Island		Yuncker, T. G. (1945) (p. 41) Vouchers cited: Yuncker 9083 (BISH), Garber 603 (BISH), Garber 634 (BISH), Garber 645a (BISH)
American Samoa Manu‘a Islands	Ta‘u Island	introduced	Ragone, Diane/Lorence, David H. (2003) (p. 49)
American Samoa Tutuila Islands	Tutuila Island		Swarbrick, John T. (1997) (p. 91)
American Samoa Tutuila Islands	Tutuila Island	introduced abo. intro.	Whistler, W. A. (1980) (p. 131)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Northern Mariana Islands		Swarbrick, John T. (1997) (p. 91)
Cook Islands Southern Cook Islands	Mangaia Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 16)
Cook Islands Southern Cook Islands	Ma‘uke Island		McCormack, Gerald (2011) ?
Cook Islands Southern Cook Islands	Ma‘uke Island	introduced invasive	Space, James C./Flynn, Tim (2002) (p. 16) Voucher: Flynn 7086 (PTBG, BISH, NY, US)
Cook Islands Southern Cook Islands	Ma‘uke Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 385) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Federated States of Micronesia Chuuk Islands	Weno (Moen) Island	native	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109) var. strigosum f. substrigosum
Federated States of Micronesia Kosrae Island	Kosrae Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2005) (voucher ID: PTBG 42612) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	native	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109) var. strigosum f. substrigosum
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		Glassman, S. F. (1952) (p. 76) Vouchers cited: Glassman 2316 (US), Glassman 2445 (US), Kanehira 1574 (NY)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island	native	Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 92) Vouchers cited: M.V.C. Falanruw MVCF3223 (BISH), M.V.C. Falanruw MVCF5405 (BISH), R. Gallen 38 (NY, PTBG), S.F. Glassman 2316 (BISH), S.F. Glassman 2445 (BISH), E.Y. Hosokawa 3502 (BISH), T. Hosokawa 5830 (BISH), T. Hosokawa 9569 (BISH), R. Kanehira 1574 (BISH), S. Perlman 20295 (PTBG), P. Raynor 158 (NY, PTBG), M. Takamatsu 740 (BSIH)
Federated States of Micronesia Pohnpei Islands	Pohnpei Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1985) (voucher ID: PTBG 23357) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Federated States of Micronesia Yap Islands	Yap (Waqab) Island	native	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109) var. strigosum f. substrigosum
Fiji Fiji Islands	Koro Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Voucher cited: DA 11803
Fiji Fiji Islands	Matuku Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Voucher cited: Bryan 260
Fiji Fiji Islands	Mbengga Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Voucher cited: DA 9619
Fiji Fiji Islands	Ngau Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Voucher cited: Milne 150
Fiji Fiji Islands	Ovalau Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Voucher cited: Seemann 111
Fiji Fiji Islands	Taveuni Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Vouchers cited: DA 2564, Gillespie 4774
Fiji Fiji Islands	Vanua Levu Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Vouchers cited: Smith 6811, DA 11779, DA 13137
Fiji Fiji Islands	Viti Levu Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Vouchers cited: Greenwood 452, DA 513, DA 10005, B. & H. Parham 35, DA 1052
Fiji Fiji Islands	Wakaya Island	introduced invasive	Smith, Albert C. (1985) (pp. 192-193) var. strigosum. Voucher cited: Milne 377
French Polynesia Gambier Islands	Akamaru Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Gambier Islands	Akamaru Island	native	St. John, Harold (1988) (p. 15) Voucher cited: St. John 14698 (BISH)
French Polynesia Gambier Islands	Mangareva (Gambier) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen. Voucher cited: B. Huguenin s.n. (PAP) Adventice
French Polynesia Gambier Islands	Mangareva (Gambier) Island	native	St. John, Harold (1988) (p. 15) Vouchers cited: Agassiz 105 (GH), Fosberg 11065 (BISH), Fosberg 11098 (BISH), Fosberg 11117 (BISH), St. John 14482 (BISH)
French Polynesia Gambier Islands	Taravai Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Gambier Islands	Taravai Island	native	St. John, Harold (1988) (p. 15) Voucher cited: St. John 14779 (BISH)
French Polynesia Marquesas Islands	Fatu Hiva Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Marquesas Islands	Hatuta'a (Hatutu) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008) Var. strigosum Meeuwen
French Polynesia Marquesas Islands	Hiva Oa Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008) Var. strigosum Meeuwen
French Polynesia Marquesas Islands	Nuku Hiva (Nukahiva) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence 7288 (PAP) Adventice
French Polynesia Marquesas Islands	Tahuata (Tuhuata) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Lorence, David H./Wagner, Warren L. (2008) Var. strigosum Meeuwen
French Polynesia Marquesas Islands	Ua Huka (Huahuna, Uahuka) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Marquesas Islands	Ua Pou (Huapu, Uapou, Uapu) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Society Islands	Bora Bora Island	introduced	Welsh, S. L. (1998) (p. 149) Voucher cited: Grant 5026
French Polynesia Society Islands	Bora Bora Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Society Islands	Huahine Island	introduced	Welsh, S. L. (1998) (p. 149) Voucher cited: D. Bare 9
French Polynesia Society Islands	Huahine Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: D. Baré 8 (PAP) Adventice
French Polynesia Society Islands	Maupiti (Maurua) Island	introduced	Fosberg, F. R./Sachet, M.-H. (1987) (p. 35) Not seen in 1985. Voucher cited: Raynal 17860.
French Polynesia Society Islands	Mehetia (Mahetia, Meetia) Island	introduced	Welsh, S. L. (1998) (p. 149) Voucher cited: Grant 4744
French Polynesia Society Islands	Mehetia (Mahetia, Meetia) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Society Islands	Moorea Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Society Islands	Raiatea (Havai) Island	introduced	Welsh, S. L. (1998) (p. 149) Vouchers cited: Moore 3, BRY 25296, BRY 26701
French Polynesia Society Islands	Raiatea (Havai) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen. Voucher cited: J. Florence 11857 (PAP) Adventice
French Polynesia Society Islands	Tahiti Island	introduced	Welsh, S. L. (1998) (p. 149) Voucher cited: Florence 8064
French Polynesia Society Islands	Tahiti Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen. Vouchers cited: J. Florence 8064 (PAP), J. Florence 9893 (PAP), J. Florence & S. Jourdan 10719 (PAP) Adventice
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Austral (Tubuai) Islands	Raivavae (Raevavae) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Austral (Tubuai) Islands	Rimatara (Rimitara) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Austral (Tubuai) Islands	Rurutu Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
French Polynesia Austral (Tubuai) Islands	Tubuai Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) var. strigosum M.S.K. Meeuwen Adventice
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced	Wester, Lyndon (1992) (p. 138)
Japan (offshore islands) Bonin (Ogasawara) Islands	Bonin (Ogasawara) Islands		Kato, Hidetoshi (2007)
Kiribati Kiribati	Kiribati (Republic of)	introduced	Fosberg, F. R./Sachet, M.-H. (1987) (p. 16)
Marshall Islands Marshall Islands	Marshall Islands		Swarbrick, John T. (1997) (p. 91)
New Caledonia New Caledonia	New Caledonia Islands	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Niue Niue	Niue Island	introduced	Sykes, W. R. (1970) (pp. 151-152) Voucher cited: CHR 150536
Niue Niue	Niue Island	introduced invasive abo. intro.	Whistler, W. A. (1988) (p. 23)
Palau Palau (main island group)	Babeldaob Island	native	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109) var. strigosum f. strigosum.
Palau Palau (main island group)	Babeldaob Island	native	Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. Voucher specimen) var. strigosum. Voucher: BMW 6557 (BISH, BRI)
Palau Palau (main island group)	Babeldaob Island	native	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. Field notes and voucher.) var. strigosum. Voucher: Lorence 9911 (PTBG, BNM, NY, US)
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1966) (voucher ID: BNM 455) Taxon name on voucher: Desmodium heterocarpon var heterocarpon (Willd.) DC.
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1981) (voucher ID: BNM 3149) Taxon name on voucher: Desmodium heterocarpon (Willd.) DC.
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (2007) (voucher ID: BNM 4401) Taxon name on voucher: Desmodium heterocarpon (L.) DC. var. strigosum v. Meeuwen
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1991) (voucher ID: BNM 9569) Taxon name on voucher: Desmodium heterocarpon var. strigosum V. Meeuwen.
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1981) (voucher ID: BNM 3098) Taxon name on voucher: Desmodium heterocarpon var. strigosum v. Meeuwen
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1978) (voucher ID: BNM 1255) Taxon name on voucher: Desmodium heterocarpon var. strigosum v. Meeuwen
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1982) (voucher ID: BNM 3195) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau Palau (main island group)	Babeldaob Island		Beleu National Museum (Republic of Palau. Koror.) (1978) (voucher ID: BNM 10153) Taxon name on voucher: Desmodium heterocarpon cf. var. strigosum Meeuw.
Palau Palau (main island group)	Babeldaob Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 204) Taxon name on voucher: Desmodium heterocarpon (L.) DC.
Palau Palau (main island group)	Babeldaob Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1966) (voucher ID: PTBG 22484) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau Palau (main island group)	Babeldaob Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1983) (voucher ID: PTBG 22783) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau Palau (main island group)	Babeldaob Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1981) (voucher ID: PTBG 22784) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau Palau (main island group)	Koror Island	native	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109) var. strigosum f. strigosum.
Palau Palau (main island group)	Koror Island		Beleu National Museum (Republic of Palau. Koror.) (1967) (voucher ID: BNM 456) Taxon name on voucher: Desmodium heterocarpon
Palau Palau (main island group)	Koror Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1965) (voucher ID: PTBG 22782) Taxon name on voucher: Desmodium heterocarpon var. strigosum
Palau Palau (main island group)	Ngerkebesang Island	native	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 109) var. strigosum f. strigosum.
Papua New Guinea Bismarck Archipelago	Bismarck Archipelago	native	ILDIS Co-ordinating Centre (2011)
Papua New Guinea Bismarck Archipelago	Bismarck Archipelago		Peekel, P. G. [translated by E. E. Henty] (1984) (p. 237) Frequent, in grassy places, at roadsides and in scrub.
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Philippines Philippine Islands	Philippine Islands	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Philippines Philippine Islands	Philippine Islands		Merrill, Elmer D. (1923) (pp. 285-286) In thickets, open grasslands, etc., at low and medium altitudes; common.
Solomon Islands Solomon Islands	Solomon Islands	native	ILDIS Co-ordinating Centre (2011)
Solomon Islands Solomon Islands	Solomon Islands	introduced invasive	Hancock, I. R./Henderson, C. P. (1988) (p. 65)
Tonga Ha‘apai Group	Kao Island	probably introduced invasive abo. intro.	Whistler, W. A. (1992) (p. 29) Vouchers cited: Yuncker 15871, Buelow 741
Tonga Ha‘apai Group	Kao Island		Yuncker, T. G. (1959) (p. 140) Voucher cited: Yuncker 15871
Tonga Ha‘apai Group	Tofua Island	probably introduced invasive abo. intro.	Whistler, W. A. (1992) (p. 29) Vouchers cited: Hotta 4331, Scarth-Johnson 75, Buelow 599
Tonga Tonga Islands	Tonga Islands		Yuncker, T. G. (1959) (p. 140) "Frequent throughout Tonga"
Tonga Tongatapu Group	‘Eua Island	probably introduced invasive abo. intro.	Whistler, W. A. (1992) (p. 29) Vouchers cited: Lister s.n., Yuncker 15372, Sykes 655, Sykes 668, Whistler 7414
Tonga Tongatapu Group	‘Eua Island		Yuncker, T. G. (1959) (p. 140) Vouchers cited: Yuncker 15372, Yuncker 15654
Tonga Tongatapu Group	Tongatapu Island		Ellison, Joanna C. (1990) (p. 29) On outlier islet of Polo‘a.
Tonga Vava‘u Group	Late Island	probably introduced invasive abo. intro.	Whistler, W. A. (1992) (p. 29) Voucher cited: Buelow & sykes 212
Tonga Vava‘u Group	Niuafo‘ou Island	probably introduced invasive abo. intro.	Whistler, W. A. (1992) (p. 29) Vouchers cited: Yuncker 16012, Buelow 2209
Tonga Vava‘u Group	Niuafo‘ou Island		Yuncker, T. G. (1959) (p. 140) Voucher cited: Yuncker 16012
Tonga Vava‘u Group	Tafahi Island	probably introduced invasive abo. intro.	Whistler, W. A. (1992) (p. 29) Vouchers cited: Hurlimann 524, Whistler 6236
Tonga Vava‘u Group	Vava‘u Island		Yuncker, T. G. (1959) (p. 140) Cited: Crosby.
Tuvalu Tuvalu (Ellis) Islands	Tuvalu Islands		Swarbrick, John T. (1997) (p. 91)
Wallis and Futuna Wallis and Futuna (Horne) Islands	Wallis (‘Uvea) Island	introduced invasive	Meyer, Jean-Yves (2007) (p. 34)
Wallis and Futuna Wallis and Futuna (Horne) Islands	Wallis and Futuna Islands		Whistler, W. A. (1988) (p. 23)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia Australia (continental)	Queensland	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
China China	China (People's Republic of)	native	Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011) "Grasslands, grassy slopes, watersides, thickets, forests; 300-1800 m".
China China	Hong Kong	native	Wu, Te-lin (2001) (p. 143)
Colombia Colombia	Colombia (Republic of)	introduced	ILDIS Co-ordinating Centre (2011)
Indonesia Indonesia	Indonesia (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Indonesia Indonesia	West Papua Province (Indonesia) (western New Guinea Island) (formerly Irian Jaya)	native	ILDIS Co-ordinating Centre (2011)
Japan Japan	Japan (country)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Japan Ryukyu Islands	Ryukyu Islands	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Malaysia Malaysia	Malaysia (country of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Perú Perú	Perú (Republic of)	native	ILDIS Co-ordinating Centre (2011)
Singapore Singapore	Singapore (Republic of)	uncertain if introduced invasive	Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 33) Weed of uncertain origin
Taiwan Taiwan	Taiwan Island	native	Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011) "Grasslands, grassy slopes, watersides, thickets, forests; 300-1800 m".
Thailand Thailand	Thailand (Kingdom of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Vietnam Vietnam	Vietnam (Socialist Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)

Indian Ocean
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Seychelles Seychelles Islands	Seychelles Islands	native	ILDIS Co-ordinating Centre (2011)

Comments:  Cook Islands subsp. heterocarpon, all other locations var. strigosum.

Previously collected in Hawai‘i but present status unknown (Wagner et al., 1999; p. 667).

Additional information:
Additional online information about Desmodium heterocarpon is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Desmodium heterocarpon as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Desmodium heterocarpon may be available from the Germplasm Resources Information Network (GRIN).

References:

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