Chloris virgata Sw., Poaceae

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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Common name(s): [more details]

Chinese: hu wei cao

English: feather finger grass, feather windmill grass, feathered chloris, feathertop Rhodes grass, feathery Rhodes grass, oldland grass, showy chloris, sweet grass

Kosraean: muh-shah-shruh

Spanish: verdillo plumerito, zacate pluma

Habit:  grass

Description:  "Annuals; culms erect or geniculately ascending, up to 10 dm tall, occasionally rooting at lower nodes.  Basal sheaths strongly keeled and often flabellate, glabrous; ligule inconspicuous, membranous, ca. 0.5 mm long; blades flat, 10-30 cm long, 2-6 mm wide, tapering at apex, upper surface scabrous, lower surface glabrous.  Inflorescences of 4-12 digitate, erect to spreading, white to yellowish brown, feathery spikes 2-10 cm long, axils silky villous; spikelets crowded, (2) 3-flowered; first glume 1.5-2.5 mm long, second glume 2.5-4.5 mm long, apex more or less with a short awn-point; first lemma obliquely obovate in side view, 2.5-4 mm long, margins and keel ciliate, apex with a crown of hairs 1.5-4 mm long, awn 5-15 mm long, callus rounded, pilose; palea lanceolate, ca. 0.3 mm long, 2-nerved, apical margin ciliate; second lemma slightly projecting from the side of the fertile lemma, oblong, 2-2.5 mm long, glabrous, with an awn 5-12 mm long; apical lemma an awnless clavate scale 0.5-1.2 mm long.  Caryopsis golden, cylindrical, 1.5-2 mm long."  (Wagner et al., 1999; pp. 1515-1516).

Description from GrassBase.

Habitat/ecology:  Aggressive invader of degraded land that out-competes native species (Smith, 2002; p. 72).  In Hawai‘i, "naturalized and common along roadsides, in dry pastures, and in open, mesic shrubland, 0-410 m". (Wagner et al., 1999; pp. 1515-1516). In New Caledonia, "assez commun en terrains vagues près de Nouméa" (MacKee, 1994; p. 58). Arid lowlands in the Galápagos Islands (McMullen, 1999; p. 240). In Australia, "a variety of habitats, particularly disturbed areas and roadsides; also levee banks, creek lines and gardens" (Smith, 2002; p. 72).

Propagation:  "Seed is dispersed short distance by wind and water, also as a contaminant of hay and by animals".

Native range:  Neotropics

Presence:

Pacific
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Saipan Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 34)
Commonwealth of the Northern Mariana Islands (US) Northern Mariana Islands	Tinian Island	introduced	Raulerson, L. (2006) (p. 7)
Ecuador (Galápagos Islands) Española Group	Española Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Floreana Group	Campéon Islet, Floreana Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Floreana Group	Floreana Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Floreana Group	Gardner por Floreana Island, Floreana Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Cruz Group	Baltra Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Cruz Group	Daphne Mayor Island, Santa Cruz Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Cruz Group	Santa Cruz Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Cruz Group	Seymour Norte Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santa Fé Group	Santa Fé Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santiago Group	Rábida Island	possibly introduced invasive	Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands) Santiago Group	Santiago Island	possibly introduced invasive	Charles Darwin Foundation (2008)
French Polynesia Society Islands	Tahiti Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Voucher cited: J. Florence 10717 (PAP) Adventice
French Polynesia Tuamotu Archipelago	Makatea (Ma‘atea) Island	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
French Polynesia Tuamotu Archipelago	Tikehau (Tikahau) Atoll	introduced invasive	Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011) Adventice
State of Hawaii Hawaiian Islands	Hawai‘i (Big) Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516)
State of Hawaii Hawaiian Islands	Kaho‘olawe Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516)
State of Hawaii Hawaiian Islands	Kaua‘i Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516)
State of Hawaii Hawaiian Islands	Kure Atoll	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516)
State of Hawaii Hawaiian Islands	Kure Atoll	introduced	Starr, Forest/Martz, Kim/Loope, Lloyd (2001) (p. 10) Previously reported, not observed in this survey.
State of Hawaii Hawaiian Islands	Lāna‘i Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516)
State of Hawaii Hawaiian Islands	Ni‘ihau, Lehua Islet	introduced invasive	Wood, K. R./LeGrande, Maya (2006) (pp. 25-26) Voucher cited: K.R. Wood 11538 (PTBG)
State of Hawaii Hawaiian Islands	Maui Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516)
State of Hawaii Hawaiian Islands	Moloka‘i Island	introduced invasive	Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 1515-1516) Voucher cited: Munro 127 (BISH)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive	Herbst, Derral R./Clayton, W. D. (1998) (p. 21) Voucher cited: Uta‘i 8 (BISH)
State of Hawaii Hawaiian Islands	O‘ahu Island	introduced invasive	Imada, Clyde T./Staples, George W./Herbst, Derral R. (2000) (pp. 14-15) Voucher cited: J. Rémy 26 (BISH)
Nauru Nauru Island	Nauru Island	introduced	Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 34)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	Tassin, Jacques (2005)
New Caledonia New Caledonia Archipelago	Île Grande Terre	introduced invasive	MacKee, H. S. (1994) (p. 58) Voucher cited: Guillaumin & Baumann 6822
Palau Palau (main island group)	Babeldaob Island	introduced invasive	Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 4) Voucher: Lorence 9816 (PTBG, BNM, BISH, NY, US)
Palau Palau (main island group)	Babeldaob Island		National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 109) Taxon name on voucher: Chloris virgata Sw.
Papua New Guinea Papua New Guinea (eastern New Guinea Island)	Papua New Guinea (eastern New Guinea Island)		Henty, E. E. (1969) (p. 50)
United States (other Pacific offshore islands) Midway Atoll	Midway Atoll	introduced invasive	Starr, Forest/Martz, Kim (2000) (p. 11) Voucher cited: Starr & Martz 990507-5 (BISH)
Vanuatu New Hebrides Islands	Vanuatu (Republic of)		Swarbrick, John T. (1997) (p. 89)

Pacific Rim
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
Australia Australia (continental)	Northern Territory	introduced invasive	Smith, Nicholas M. (2002) (p. 72)
Australia Australia (continental)	Queensland	introduced invasive	Smith, Nicholas M. (2002) (p. 72)
Chile (continental) Chile	Chile (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
China China	China (People's Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Colombia Colombia	Colombia (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Ecuador (Mainland) Ecuador	Ecuador (Republic of) (continental)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
El Salvador El Salvador	El Salvador (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Guatemala Guatemala	Guatemala (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Honduras Honduras	Honduras (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Indonesia Indonesia	Indonesia (Republic of)		Hafliger, Ernst/Scholz, Hildemar (1980) (p. 52)
Japan Japan	Japan (country)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Japan Japan	Japan (country)	introduced	Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 192)
South Korea South Korea	South Korea (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Mexico Mexico	Mexico (United Mexican States)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Nicaragua Nicaragua	Nicaragua (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Panama Panama	Panama (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)
Perú Perú	Perú (Republic of)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)

Also reported from
Country/Terr./St. & Island group	Location	Cited status & Cited as invasive & Cited as cultivated & Cited as aboriginal introduction?	Reference & Comments
United States (continental except west coast) United States (other states)	United States (other states)	native	U.S. Dept. Agr., Agr. Res. Serv. (2011)

Control: 

Physical:  Pull or dig out.

Chemical:  Spray with herbicide (Weber, 2003; p. 103).

Additional information:
Additional online information about Chloris virgata is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Chloris virgata as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Chloris virgata may be available from the Germplasm Resources Information Network (GRIN).

References:

Bishop Museum (Honolulu). 1949. Voucher specimen #BISH 123718 (Glassman, S.F. 2668).

Charles Darwin Foundation. 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Charles Darwin Foundation, Galapagos, Ecuador.

Charles Darwin Research Station. 2005. CDRS Herbarium records.

Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. 2011. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP). (online resource).

Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1987. A geographical checklist of the Micronesian monocotyledonae. Micronesica 20:1-126.

Hafliger, Ernst/Scholz, Hildemar. 1980. Grass weeds, vol. 2. CIBA-GEIGY Ltd., Basle, Switzerland. 137 pp. + plates.

Henty, E. E. 1969. A manual of the grasses of New Guinea. Department of Forests, Division of Botany, Botany Bull. No. 1. Lae, New Guinea. 215 pp.

Herbst, Derral R./Clayton, W. D. 1998. Notes on the grasses of Hawai‘i: new records, corrections, and name changes. In: Evenhuis, Neal L. and Miller, Scott, E., eds. Records of the Hawaii Biological Survey for 1997. Part 1: Articles. Bishop Museum Occasional Papers. 55:17-38.

Imada, Clyde T./Staples, George W./Herbst, Derral R. 2000. New Hawaiian plant records for 1999. In: Evenhuis, Neal L. and Eldredge, Lucius G., eds. Records of the Hawaii Biological Survey for 1999. Part 1: Articles. Bishop Museum Occasional Papers. 63:9-16.

MacKee, H. S. 1994. Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie. Muséum National d'Histoire Naturelle, Paris, 164 p.

McMullen, C. K. 1999. Flowering plants of the Galápagos. Comstock Pub. Assoc., Ithaca, N.Y. 370 p.

Mito, Toshikazu/Uesugi, Tetsuro. 2004. Invasive alien species in Japan: the status quo and the new regulation for prevention of their adverse effects. Global Environmental Research 8(2)/2004: 171-191.

National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.). 2008. Voucher specimen #PTBG109(David H. Lorence 9816).

Raulerson, L. 2006. Checklist of Plants of the Mariana Islands. University of Guam Herbarium Contribution 37:1-69. .

Smith, Nicholas M. 2002. Weeds of the wet/dry tropics of Australia - a field guide. Environment Centre NT, Inc. 112 pp.

Space, James C./Lorence, David H./LaRosa, Anne Marie. 2009. Report to the Republic of Palau: 2008 update on Invasive Plant Species. USDA Forest Service, Hilo, Hawai‘i. 227 pp.

Starr, Forest/Martz, Kim. 2000. New plant records from Midway Atoll for 1999. In: Evenhuis, Neal L. and Eldredge, Lucius G., eds. Records of the Hawaii Biological Survey for 1999. Part 2: Notes. Bishop Museum Occasional Papers. 64:10-12.

Starr, Forest/Martz, Kim/Loope, Lloyd. 2001. Botanical inventory of Kure Atoll. United States Geological Survey, Biological Resources Division, Haleakala Field Station, Hawai‘i. 22 pp.

Swarbrick, John T. 1997. Weeds of the Pacific Islands. Technical paper no. 209. South Pacific Commission, Noumea, New Caledonia. 124 pp.

Tassin, Jacques. 2005. Jacques Tassin (IAC-CIRAD), personal communication.

U.S. Dept. Agr., Agr. Res. Serv. 2011. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.

Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawai‘i Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).

Weber, Ewald. 2003. Invasive plants of the World. CABI Publishing, CAB International, Wallingford, UK. 548 pp.

Wiggins, I. L./Porter, D. M. 1971. Flora of the Galapágos Islands. Stanford University Press. 998 pp.

Wood, K. R./LeGrande, Maya. 2006. An annotated checklist and new island records of flowering plants from Lehua Islet, Ni‘ihau, Hawai‘i. In: Evenhuis, Neal L. and Eldredge, Lucias G., eds. Records of the Hawaii Biological Survey for 2004-2005. Part 1: Articles. Bishop Museum Occasional Papers 87:19-29.

Zhengyi, Wu/Raven, Peter H./Deyuan, Hong. 2011. Flora of China (online resource).