|
L., Amaranthaceae |
|
Present on Pacific Islands? yes
Primarily a threat at high elevations? no
Other Latin names: Amaranthus gracilis Desf., nom. nud.
Common name(s): [more details]
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Chamorro: kuletes apaka |
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Chinese: zhou guo xian |
|
English: green amaranth, pigweed, Prince of Wales feather, slender amaranth, tropical green amaranth |
|
Fijian: moca, ndriti, ngasau ni vuaka, tubua, tumbua |
|
French: amarante verte, brede malabar, fleur de jalousie |
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Hawaiian: aheahea, pakai, pākaikai, pakapakai |
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Hindi: chauraiya, chauraiya, choroiya |
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I-Kiribati: te moota, te mota |
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Japanese: aobiyu, honaga-inubiyu |
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Maori (Cook Islands): puekao, vaine ara |
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Marquesan: puaio, upootii |
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Niuean: katule |
|
Spanish: amarantus, bledo, bredo blanco |
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Tahitian: ea ea mata, huinihera, upooti |
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Tongan: longolongouha, tupua |
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Ulithian: waluwol |
Habit: herb
Description: "Monoecious annual herbs; stems erect or occasionally ascending, 1-8 (-10) dm long, sparingly to densely branched, striate, glabrous and usually becoming pubescent with multicellular hairs above. Leaves deltate-ovate to narrowly rhombic, blades 2-7 cm long, 1.5-5.5 cm wide, glabrous or lower surface pilose along the veins, apex usually narrow and with a small narrow emargination, petioles 1-10 cm long. Flowers green, in slender, axillary or terminal and often paniculate spikes, sometimes in axillary clusters in lower part of plant, both sexes mixed throughout the spikes, but pistillate flowers more numerous, bracts and bracteoles whitish, deltate-ovate to broadly lanceolate, membranous, with a short, pale or reddish awn; sepals 3 (4), those of staminate flowers ovate-oblong, ca. 1.5 mm long, apex acute, mucronate, those of pistillate flowers narrowly spatulate to oblong, 1.3-1.8 mm long, apex more or less mucronate; stigmas 2-3. Fruit subglobose, 1.3-1.5 mm long, not or only slightly exceeding the sepals, indehiscent or rupturing irregularly at maturity, conspicuously rugose throughout. Seeds dark brown to black, more or less shiny, slightly compressed, 1-1.3 mm long, reticulate and with shallow outgrowths on the reticulum" (Wagner et al., 1999; p. 189).
Habitat/ecology: "A weed, but rarely also cultivated (for edible leaves); often prostrate, flattened; waysides, vacant lots, crevices of sidewalks, edge of asphalt strips, etc." (Stone, 1970; p. 269). "Occasional in croplands and waste places." (Whistler, 1988; p. 7).
In Hawaii, "the most common naturalized species of Amaranthus, occurring in low elevation, disturbed habitats" (Wagner et al., 1999; p. 189). In Fiji, "occurring near sea level as a naturalized weed of cultivation and waste places, often locally abundant, common along roadsides and in gardens, plantations and canefields" (Smith, 1981; p. 286). On Niue, "a common weed around houses and in plantations" (Sykes, 1970; p. 41). In Tonga, "a weed of plantations and waste areas" (Yuncker, 1959; p. 107). In New Zealand, "crops, sand dunes" (Webb et al., 1988; p. 108). In Papua New Guinea, "occurring as a weed in gardens and waste places, also in secondary vegetation, at 0-2000 m altitude" (Womersley, 1978; p. 30).
Propagation: Seed
Native range: "Origin obscure though possibly South America" (GRIN)
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
American Samoa
Manua Islands |
Olosega Island |
Yuncker, T. G. (1945) (p. 37)
Foucher cited: Garber 1069 (BISH) |
|
|
American Samoa
Manua Islands |
Tau Island |
Whistler, W. A. (1990)
Vouchers cited: Whistler 1374 |
|
|
American Samoa
Manua Islands |
Tau Island |
Yuncker, T. G. (1945) (p. 37)
Vouchers cited: Yuncher 9197 (BISH), W. Harris 9 (BISH), W. Harris 262 (BISH) |
|
|
American Samoa
Tutuila Islands |
Tutuila Island | Swarbrick, John T. (1997) (p. 88) | |
|
American Samoa
Tutuila Islands |
Tutuila Island | Anon. (1988) (p. 1) | |
|
American Samoa
Tutuila Islands |
Tutuila Island |
introduced
invasive |
Setchell, William Albert (1924) (p. 96) |
|
Australia (Pacific offshore islands)
Lord Howe Island |
Lord Howe Island |
introduced
|
Pickard, J. (1984) (p. 205) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Agrigan (Agrihan) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Agrigan (Agrihan) Island |
introduced
invasive |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 23)
Voucher cited: Fosberg 2160 (US) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Aguijan (Agiguan, Aguihan) Island |
introduced
|
Raulerson, L. (2006) (p. 29) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Alamagan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Alamagan Island |
introduced
invasive |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 23)
Vouchers cited: Falanruw 1947 (US), Fosberg 31653 (US) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Anatahan Island |
introduced
|
Raulerson, L. (2006) (p. 29) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Pagan Island |
introduced
|
Raulerson, L. (2006) (p. 29) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Pagan Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1981) (voucher ID: PTBG 26474)
Taxon name on voucher: Amaranthus viridis |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1966) (voucher ID: PTBG 26472)
Taxon name on voucher: Amaranthus viridis |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
introduced
invasive |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 23)
Vouchers cited: Falanruw 1732 (US), Falanruw 1786 (US), Evans 2358 (US, BISH, Fo, NY), Falanruw 1745 (US) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1985) (voucher ID: PTBG 26470)
Taxon name on voucher: Amaranthus viridis |
|
|
Cook Islands
Southern Cook Islands |
Aitutaki Atoll | McCormack, Gerald (2011) | |
|
Cook Islands
Southern Cook Islands |
Atiu Island | McCormack, Gerald (2011) | |
|
Cook Islands
Southern Cook Islands |
Mangaia Island | McCormack, Gerald (2011) | |
|
Cook Islands
Southern Cook Islands |
Mitiaro Island | McCormack, Gerald (2011) | |
|
Cook Islands
Southern Cook Islands |
Rarotonga Island | McCormack, Gerald (2011) | |
|
Ecuador (Galápagos Islands)
Isabela Group |
Isabela Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Isabela Group |
Volcán Sierra Negra, Isabela Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Isabela Group |
Volcán Wolf, Isabela Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Pinta Group |
Pinta Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
San Cristóbal Group |
San Cristóbal Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Santa Cruz Group |
Santa Cruz Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Santa Fé Group |
Santa Fé Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Ecuador (Galápagos Islands)
Santiago Group |
Santiago Island |
introduced
invasive |
Charles Darwin Foundation (2008) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane/Waguk, Erick E. (2000) (p. Voucher specimen)
Voucher: BMW 5779 (BISH, CANB) |
|
Federated States of Micronesia
Kosrae Island |
Kosrae Island |
introduced
invasive |
Lorence, David H./Flynn, Timothy (2010) (p. 12)
Naturalized |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
invasive |
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 66)
Voucher cited: M. Takamatsu 1018 (BISH) Naturalized |
|
Federated States of Micronesia
Yap Islands |
Fais Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Federated States of Micronesia
Yap Islands |
Fais Island |
native
|
Fosberg, F. R./Evans, M. (1969) (p. 6)
Voucher cited: Evans 343 (US) |
|
Federated States of Micronesia
Yap Islands |
Satawal Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Federated States of Micronesia
Yap Islands |
Satawal Island |
introduced
invasive |
Fosberg, F. R. (1969) (p. 7)
Voucher cited: Fosberg 46888a (US) |
|
Federated States of Micronesia
Yap Islands |
Ulithi Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Federated States of Micronesia
Yap Islands |
Yap (Waqab) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Fiji
Fiji Islands |
Moala Island |
introduced
invasive |
Smith, Albert C. (1981) (p. 286)
As A. gracilis. Voucher cited: Milne 111 |
|
Fiji
Fiji Islands |
Taveuni Island |
introduced
invasive |
Smith, Albert C. (1981) (p. 286)
As A. gracilis. Vouchers cited: DA 8878, DA 5743 |
|
Fiji
Fiji Islands |
Vanua Levu Island |
introduced
invasive |
Smith, Albert C. (1981) (p. 286)
As A. gracilis. Voucher cited: DA 8952 |
|
Fiji
Fiji Islands |
Viti Levu Island |
introduced
invasive |
Smith, Albert C. (1981) (p. 286)
As A. gracilis. |
|
French Polynesia
Gambier Islands |
Mangareva (Gambier) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Gambier Islands |
Mangareva (Gambier) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Gambier Islands |
Mangareva (Gambier) Island |
introduced
|
St. John, Harold (1988) (p. 13)
Vouchers cited: Agassiz 96 (GH), Fosberg 11091 (BISH) |
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Marquesas Islands |
Fatu Hiva Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39578)
Taxon name on voucher: Amaranthus viridis |
|
|
French Polynesia
Marquesas Islands |
Fatu Huku Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: R.L. Oliver & P.A. Schäfer 3100b (PAP) Adventice |
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 41673)
Taxon name on voucher: Amaranthus viridis |
|
|
French Polynesia
Marquesas Islands |
Hiva Oa Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39579)
Taxon name on voucher: Amaranthus viridis |
|
|
French Polynesia
Marquesas Islands |
Mohotani (Mohotane, Mohotane) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Mohotani (Mohotane, Mohotane) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Mohotani (Mohotane, Mohotane) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Nuku Hiva (Nukahiva) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Marquesas Islands |
Tahuata (Tuhuata) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Tahuata (Tuhuata) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Tahuata (Tuhuata) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Vouchers cited: K.R. Wood, S.P. Perlman, J.-Y. Meyer & L. Teatiu 6407 (PAP), S.P. Perlman, K.R. Wood & J.-Y. Meyer 15862 (PAP) Adventice |
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 26262)
Taxon name on voucher: Amaranthus viridis L. |
|
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1997) (voucher ID: PTBG 28578)
Taxon name on voucher: Amaranthus viridis L. |
|
|
French Polynesia
Marquesas Islands |
Ua Huka (Huahuna, Uahuka) Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 40872)
Taxon name on voucher: Amaranthus viridis |
|
|
French Polynesia
Marquesas Islands |
Ua Pou (Huapu, Uapou, Uapu) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Marquesas Islands |
Ua Pou (Huapu, Uapou, Uapu) Island |
introduced
invasive |
Lorence, David H./Wagner, Warren L. (2008) |
|
French Polynesia
Marquesas Islands |
Ua Pou (Huapu, Uapou, Uapu) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Society Islands |
Huahine Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 27)
Voucher cited: Bare 117 |
|
French Polynesia
Society Islands |
Huahine Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Society Islands |
Huahine Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Society Islands |
Moorea Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Society Islands |
Mopelia (Mapetia, Maupihaa, Maupihoa, Mopihaa) Atoll |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Society Islands |
Mopelia (Mapetia, Maupihaa, Maupihoa, Mopihaa) Atoll |
introduced
invasive |
Sachet, M.-H. (1983) (p. 26)
Voucher cited: Sachet 966 (US) |
|
French Polynesia
Society Islands |
Mopelia (Mapetia, Maupihaa, Maupihoa, Mopihaa) Atoll |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 27)
Voucher cited: BRY 26491 |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Society Islands |
Raiatea (Havai) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive |
Welsh, S. L. (1998) (p. 27)
Vouchers cited: BRY 25456, BRY 25998 |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Society Islands |
Tahiti Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Tuamotu Archipelago |
Raroia Atoll |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Tuamotu Archipelago |
Raroia Atoll |
introduced
|
Doty, Maxwell S. (1954) (p. 36) |
|
French Polynesia
Tuamotu Archipelago |
Raroia Atoll |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Tuamotu Archipelago |
Takapoto Atoll |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Tuamotu Archipelago |
Takapoto Atoll |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Tuamotu Archipelago |
Toau (Tovau) Atoll |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 10242 (PAP) Adventice |
|
French Polynesia
Austral (Tubuai) Islands |
Rapa Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Austral (Tubuai) Islands |
Rapa Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: N. Hallé 7687 (PAP) Adventice |
|
French Polynesia
Austral (Tubuai) Islands |
Rimatara (Rimitara) Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Austral (Tubuai) Islands |
Rimatara (Rimitara) Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
French Polynesia
Austral (Tubuai) Islands |
Rurutu Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Austral (Tubuai) Islands |
Rurutu Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 9175 (PAP) Adventice |
|
French Polynesia
Austral (Tubuai) Islands |
Tubuai Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
French Polynesia
Austral (Tubuai) Islands |
Tubuai Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Adventice |
|
Guam
Guam Island |
Guam Island |
introduced
invasive |
Stone, Benjamin C. (1970) (p. 269) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Guam
Guam Island |
Guam Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1966) (voucher ID: PTBG 26473)
Taxon name on voucher: Amaranthus viridis |
|
|
State of Hawaii
Hawaiian Islands |
Hawaii (Big) Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
State of Hawaii
Hawaiian Islands |
Kahoolawe Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
State of Hawaii
Hawaiian Islands |
Kure Atoll |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
State of Hawaii
Hawaiian Islands |
Kure Atoll |
introduced
invasive |
Starr, Forest/Martz, Kim/Loope, Lloyd (2001) (p. 8)
Uncommon near quarters. |
|
State of Hawaii
Hawaiian Islands |
Lānai Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
State of Hawaii
Hawaiian Islands |
Laysan Island |
introduced
invasive |
Shannon, Robynn K./Wagner, Warren L. (1996) (p. 13)
Voucher cited: Snyder s.n. (US) |
|
State of Hawaii
Hawaiian Islands |
Maui Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
State of Hawaii
Hawaiian Islands |
Molokai Island |
introduced
invasive |
Hughes, Guy D'Oyly (1995) (pp. 1-2)
Voucher cited: Hughes s.n. (BISH) |
|
State of Hawaii
Hawaiian Islands |
Niihau Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R. (1995) (p. 14) |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 189) |
|
Japan (offshore islands)
Bonin (Ogasawara) Islands |
Bonin (Ogasawara) Islands |
introduced
|
Kato, Hidetoshi (2007) |
|
Kiribati
Line Islands |
Malden Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 37227)
Taxon name on voucher: Amaranthus viridis L. |
|
|
Kiribati
Line Islands |
Teraina (Washington) Island |
introduced
invasive |
Wester, Lyndon (1985) (p. 21)
Uncommon. |
|
Kiribati
Line Islands |
Teraina (Washington) Island |
introduced
invasive |
Christophersen, Erling (1927) (pp. 55, 78)
Lawns and waste places at the settlement. voucher cited: Bergman 101 |
|
Kiribati
Phoenix Islands |
Kanton Island |
introduced
invasive |
Fosberg, F. R./Stoddart, D. R. (1994) (p. 8)
Rare. Voucher cited: Fosberg 55726 (US) |
|
Kiribati
Tungaru (Gilbert) Islands |
Tarawa Atoll |
Swarbrick, John T. (1997) (p. 88)
Presence not verified |
|
|
Marshall Islands
Ralik Chain |
Enewetak (Ãnewetak, Eniwetok) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Marshall Islands
Ralik Chain |
Enewetak (Ãnewetak, Eniwetok) Atoll |
introduced
invasive |
St. John, Harold (1960) (p. 326) |
|
Marshall Islands
Ralik Chain |
Enewetak (Ãnewetak, Eniwetok) Atoll |
introduced
invasive |
Fosberg, F. R. (1955) (p. 9)
Voucher cited: Fosberg 24372 (US) |
|
Marshall Islands
Ralik Chain |
Jaluit (Jãlwõj) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Marshall Islands
Ralik Chain |
Jaluit (Jãlwõj) Atoll |
introduced
invasive |
Fosberg, F. R./Sachet, M.-H. (1962) (p. 16)
Voucher cited: Fosberg 39473 (US) |
|
Marshall Islands
Ralik Chain |
Kwajalein (Kuwajleen) Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Marshall Islands
Ratak Chain |
Arno Atoll |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Marshall Islands
Ratak Chain |
Majuro (Mãjro) Atoll |
introduced
|
Fosberg, F. Raymond (1990) (p. 35) |
|
Nauru
Nauru Island |
Nauru Island |
introduced
invasive |
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (pp. 85-86)
Vouchers cited: Scully 143N (US?), Thaman & Manner 26 (SUVA) |
|
New Caledonia
Récifs D'Entrecasteaux (D'Entrecasteaux Reefs) |
Île Surprise |
introduced
invasive |
MacKee, H. S. (1994) (p. 12)
Voucher cited: Boulet in MacKee 31447 |
|
New Caledonia
Îles Loyauté (Loyalte Islands) |
Île Lifou |
introduced
invasive |
MacKee, H. S. (1994) (p. 12)
Voucher cited: MacKee 28330 |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive |
Tassin, Jacques (2005) |
|
New Caledonia
New Caledonia Archipelago |
Île Grande Terre |
introduced
invasive |
MacKee, H. S. (1994) (p. 12)
Vouchers cited: Labillardière s.n., Pancher 437, Balansa 522, Buchholz 1781, MacKee 20043, MacKee 25248, MacKee 39764 |
|
Niue
Niue |
Niue Island |
introduced
invasive |
Sykes, W. R. (1970) (p. 41)
Vouchers cited: CHR 169884, CHR 169887 |
|
Niue
Niue |
Niue Island |
introduced
invasive |
Whistler, W. A. (1988) (p. 7) |
|
Niue
Niue |
Niue Island |
introduced
invasive |
Yuncker, T. G. (1943) (p. 51)
Voucher cited: Yuncker 9866 (BISH) "A weedy plant occasional in plantations, dooryards, and waste areas". |
|
Niue
Niue |
Niue Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 38) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 100) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 69) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 96) |
|
Palau
Palau (main island group) |
Koror Island |
Beleu National Museum (Republic of Palau. Koror.) (1974) (voucher ID: BNM 10046)
Taxon name on voucher: Amaranthus viridis L. |
|
|
Palau
Palau (main island group) |
Koror Island |
Beleu National Museum (Republic of Palau. Koror.) (1969) (voucher ID: BNM 62)
Taxon name on voucher: Amaranthus viridis L. |
|
|
Palau
Palau (main island group) |
Ngercheu Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 108)
Carp Resort |
|
Palau
Palau (main island group) |
Ngercheu Island |
introduced
invasive |
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 62)
Carp Resort |
|
Palau
Palau (main island group) |
Ngerkebesang Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 96) |
|
Palau
Sonsorol Group |
Sonsorol Island |
introduced
invasive |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109) |
|
Papua New Guinea
Bismarck Archipelago |
Bismarck Archipelago |
invasive
|
Peekel, P. G. [translated by E. E. Henty] (1984) (p. 168)
Common weed in plantations, on rubbish heaps and rough lawns. |
|
Papua New Guinea
Papua New Guinea (eastern New Guinea Island) |
Papua New Guinea (eastern New Guinea Island) |
invasive
|
Henty, E. E./Pritchard, G. H. (1975) (p. 54) |
|
Papua New Guinea
Papua New Guinea (eastern New Guinea Island) |
Papua New Guinea (eastern New Guinea Island) | Waterhouse, D. F. (1997) (p. 59) | |
|
Papua New Guinea
Papua New Guinea (eastern New Guinea Island) |
Papua New Guinea (eastern New Guinea Island) |
invasive
|
Womersley, John S., ed. (1978) (pp. 29-30) |
|
Philippines
Philippine Islands |
Philippine Islands |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
|
Philippines
Philippine Islands |
Philippine Islands |
probably introduced
invasive |
Merrill, Elmer D. (1923) (pp. 128-129)
At low and medium altitudes; a common weed, probably introduced. |
|
Pitcairn Islands
Pitcairn Islands |
Pitcairn Island |
introduced
invasive |
Florence, J. (2004) (pp. 63-65) |
|
Pitcairn Islands
Pitcairn Islands |
Pitcairn Island |
introduced
invasive |
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 10750 (PAP), J. Florence 10810 (PAP) Adventice |
|
Samoa
Western Samoa Islands |
Western Samoa Islands | Whistler, W. A. (1988) (p. 7) | |
|
Solomon Islands
Solomon Islands |
Solomon Islands | Swarbrick, John T. (1997) (p. 88) | |
|
Solomon Islands
Solomon Islands |
Solomon Islands |
introduced
invasive cultivated |
Hancock, I. R./Henderson, C. P. (1988) (p. 46) |
|
Tonga
Haapai Group |
Haapai Islands |
Whistler, W. A. (1992) (p. 2)
Voucher cited: Soakai 294 |
|
|
Tonga
Tonga Islands |
Tonga Islands | Whistler, W. A. (1988) (p. 7) | |
|
Tonga
Tonga Islands |
Tonga Islands |
Yuncker, T. G. (1959) (p. 107)
"Throughout Tonga". |
|
|
Tonga
Tongatapu Group |
Eua Island |
Whistler, W. A. (1992) (p. 2)
Voucher cited: Soakai 397 |
|
|
Tonga
Tongatapu Group |
Eua Island |
Yuncker, T. G. (1959) (p. 107)
Voucher cited: Yuncker 15470 |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Soakai 440, Soakai 461, Soakai 609 |
|
|
Tonga
Tongatapu Group |
Tongatapu Island |
Yuncker, T. G. (1959) (p. 107)
Vouchers cited: Yuncker 15164, Hürlimann 100 |
|
|
Tonga
Vavau Group |
Late Island |
introduced
|
Sykes, W. R. (1981) (p. 348) |
|
Tonga
Vavau Group |
Tafahi Island |
Yuncker, T. G. (1959) (p. 107)
Voucher cited: Hürlimann 541 |
|
|
Tonga
Vavau Group |
Vavau Island |
Whistler, W. A. (1992) (p. 2)
Voucher cited: Soakai 880 |
|
|
Tuvalu
Tuvalu (Ellis) Islands |
Tuvalu Islands | Swarbrick, John T. (1997) (p. 88) | |
|
United States (other Pacific offshore islands)
Johnston Atoll |
Johnston Island |
introduced
invasive |
Amerson, A. Binion, Jr./Shelton, Philip C. (1976) (p. 55) |
|
United States (other Pacific offshore islands)
Midway Atoll |
Midway Atoll |
introduced
invasive |
Wagner, Warren L./Herbst, Derral R. (1995) (p. 14)
Vouchers cited: Herbst & Takeuchi 9074 (BISH), Herbst & Takeuchi 9080 (BISH) |
|
United States (other Pacific offshore islands)
Midway Atoll |
Sand Island |
introduced
invasive |
Starr, Forest/Starr, Kim/Loope, Lloyd (2008) (pp. B-3) |
|
United States (other Pacific offshore islands)
Wake Islands |
Wake Island |
introduced
invasive |
Fosberg, F. R. (1959) (p. 13)
Voucher cited: Fosberg 34446 |
|
United States (other Pacific offshore islands)
Wake Islands |
Wake Island |
introduced
invasive |
Fosberg, F. R./Sachet, M.-H. (1969) (p. 9)
Voucher cited: Fosberg 43517 (US) |
|
United States (other Pacific offshore islands)
Wake Islands |
Wake Island |
introduced
invasive |
Fosberg, F. R. (1955) (p. 9)
Voucher cited: Fosberg 34446 (US) |
|
Vanuatu
New Hebrides Islands |
Vanuatu (Republic of) | Swarbrick, John T. (1997) (p. 88) | |
|
Wallis and Futuna
Wallis and Futuna (Horne) Islands |
Wallis and Futuna Islands | Waterhouse, D. F. (1997) (p. 59) | |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia
Australia (continental) |
New South Wales | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Australia
Australia (continental) |
Northern Territory | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Australia
Australia (continental) |
Queensland | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Cambodia
Cambodia |
Cambodia (Kingdom of) |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
|
Chile (continental)
Chile |
Chile (Republic of) | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
China
China |
China (People's Republic of) |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
"Fields, waste places". |
|
|
China
China |
China (People's Republic of) | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
China
China |
Hong Kong |
native
|
Wu, Te-lin (2001) (p. 78) |
|
Honduras
Honduras |
Honduras (Republic of) | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (pp. 20-21) | |
|
Indonesia
Indonesia |
Indonesia (Republic of) |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
|
Japan
Japan |
Japan (country) | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Japan
Japan |
Japan (country) |
introduced
|
Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 182) |
|
South Korea
South Korea |
South Korea (Republic of) | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Malaysia
Malaysia |
Malaysia (country of) |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
|
Mexico
Mexico |
Mexico (United Mexican States) | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
New Zealand
New Zealand |
New Zealand (country) |
introduced
invasive |
Webb, C. J./Sykes, W. R./Garnock-Jones, P. J. (1988) (p. 108) |
|
Perú
Perú |
Perú (Republic of) | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Singapore
Singapore |
Singapore (Republic of) |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
|
Taiwan
Taiwan |
Taiwan Island | Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 55) | |
|
Thailand
Thailand |
Thailand (Kingdom of) |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
|
Vietnam
Vietnam |
Vietnam (Socialist Republic of) |
probably native
|
Waterhouse, D. F. (1993) (pp. 63, 76) |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Australia (Indian Ocean offshore islands)
Christmas Island Group |
Christmas Island |
native
|
Orchard, Anthony E., ed. (1993) (p. 16) |
|
British Indian Ocean Territory
Chagos Archipelago |
Diego Garcia Island |
Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 150)
"Seen by Wiehe in 1939". |
|
|
British Indian Ocean Territory
Chagos Archipelago |
Diego Garcia Island |
introduced
invasive |
Whistler, W. Arthur (1996) (p. 25) |
|
Maldives
Maldive Islands |
Malè Atoll | Fosberg, F. R. (1957) (p. 18) | |
|
Mauritius
Mautitius Islands (Mauritius and Rodrigues) |
Mauritius Island | Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (pp. 20-21) | |
Additional information:
Additional online information about Amaranthus viridis is available from the Hawaiian Ecosystems at Risk project (HEAR).
Information about Amaranthus viridis as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).
Taxonomic information about Amaranthus viridis may be available from the Germplasm Resources Information Network (GRIN).
References:
Amerson, A. Binion, Jr./Shelton, Philip C. 1976. The natural history of Johnston Atoll. Atoll Research Bulletin No. 192. Smithsonian Institution, Washington. 479 pp.
Anon. 1988. Weed list for American Samoa by cropping system. Checklist dated 07/13/88, origin unknown. 3 pp.
Beleu National Museum (Republic of Palau. Koror.). 1969. Voucher specimen #BNM62(Salsedo 180).
Beleu National Museum (Republic of Palau. Koror.). 1974. Voucher specimen #BNM10046(Otobed, D.O. P-10046).
Charles Darwin Foundation. 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Charles Darwin Foundation, Galapagos, Ecuador.
Charles Darwin Research Station. 2005. CDRS Herbarium records.
Christophersen, Erling. 1927. Vegetation of the Pacific equatorial islands. Bernice P. Bishop Museum Bulletin 44 [Whippoorwill Expedition publication 2]. Honolulu. 79 pp.
Doty, Maxwell S. 1954. Floristic and Ecological Notes on Raroia Atoll, Tuamotus. Land biota: vascular plants. Atoll Research Bulletin No. 33. Pacific Science Board, National Academy of Sciences, Washington. 41 pp.
Florence, J. 2004. Flore de la Polynésie française, Vol. 2. Paris. IRE Editions, Publications Scientifiques, Collection Faune et Flore Tropicales 41. 503 pp.
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. 2011. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP). (online resource).
Fosberg, F. R. 1955. Northern Marshall Islands Expedition, 1951-1952. Land biota: vascular plants. Atoll Research Bulletin No. 39. Pacific Science Board, National Academy of Sciences, Washington. 22 pp.
Fosberg, F. R. 1957. The Maldive Islands, Indian Ocean. Atoll Research Bulletin No. 58. Pacific Science Board, National Academy of Sciences, Washington. 37 pp.
Fosberg, F. R. 1959. Vegetation and flora of Wake Island. Atoll Research Bulletin 67. Pacific Science Board, National Academy of Sciences, Washington. 20 pp.
Fosberg, F. R. 1969. Plants of Satawal Island, Caroline Islands. Atoll Research Bulletin No. 132. Smithsonian Institution, Washington. 13 pp.
Fosberg, F. R./Evans, M. 1969. A collection of plants from Fais, Caroline Islands. Atoll Research Bulletin No. 133. Smithsonian Institution, Washington. 15 pp.
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. 1975. Vascular flora of the northern Marianas Islands. Smithsonian Contrib. Bot. 22:1-45.
Fosberg, F. R./Sachet, M.-H. 1962. Vascular plants recorded from Jaluit Atoll. Atoll Research Bulletin No. 92. Pacific Science Board, National Academy of Sciences, Washington. 39 pp.
Fosberg, F. R./Sachet, M.-H. 1969. Wake Island vegetation and flora, 1961-1963. Atoll Research Bulletin No. 123. Smithsonian Institution, Washington. 15 pp.
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.
Fosberg, F. R./Stoddart, D. R. 1994. Flora of the Phoenix Islands, central Pacific. Smithsonian Institution, Washington. Atoll Research Bulletin 393. 60 pp.
Fosberg, F. Raymond. 1990. A review of the natural history of the Marshall Islands. Atoll Research Bulletin No. 330. Smithsonian Institution, Washington. 100 pp.
Hancock, I. R./Henderson, C. P. 1988. Flora of the Solomon Islands. Research Bulletin No. 7. Ministry of Agriculture and Lands, Honiara. 203 pp.
Henty, E. E./Pritchard, G. H. 1975. Weeds of New Guinea and their control. 2nd edition. Department of Forests, Division of Botany, Botany Bull. No. 7. Lae, Papua New Guinea. 180 pp.
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. 2010. Checklist of the vascular plants of Pohnpei with local names and uses. Allertonia, in press. National Tropical Botanical Garden, Lawai, Hawaii. 146 pp.
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James. 1997. World weeds: natural histories and distribution. John Wiley & Sons. 1129 pp.
Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. 1979. A geographical atlas of world weeds. John Wiley & Sons, New York. 391 pp.
Hughes, Guy D'Oyly. 1995. New Hawaiian plant records II. In: Evenhuis, Neal L. and Miller, Scott, E., eds. Records of the Hawaii Biological Survey for 1994. Part 2: Notes. Bishop Museum Occasional Papers. 42:1-10.
Kato, Hidetoshi. 2007. Herbarium records of Makino Herbarium, Tokyo Metropolitan University. Personal communication.
Lorence, David H./Flynn, Timothy. 2010. Checklist of the plants of Kosrae. Unpublished checklist. National Tropical Botanical Garden, Lawai, Hawaii. 26 pp.
Lorence, David H./Wagner, Warren L. 2008. Flora of the Marquesas Islands. National Tropical Botanical Garden and the Smithsonian Institution. Online database.
MacKee, H. S. 1994. Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie. Muséum National d'Histoire Naturelle, Paris, 164 p.
McCormack, Gerald. 2011. Cook Islands Biodiversity Database, Version 2007.2. Cook Islands Natural Heritage Trust, Rarotonga.
Merrill, Elmer D. 1923. An enumeration of Philippine flowering plants, vol. 2 [reprint]. Bureau of Printing, Manila. 530 pp.
Mito, Toshikazu/Uesugi, Tetsuro. 2004. Invasive alien species in Japan: the status quo and the new regulation for prevention of their adverse effects. Global Environmental Research 8(2)/2004: 171-191.
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.). 1995. Voucher specimen #PTBG37227(Art Whistler W 9887).
Orchard, Anthony E., ed. 1993. Flora of Australia. Vol. 50, Oceanic islands 2. Australian Government Publishing Service, Canberra.
Parham, J. W. 1958. The weeds of Fiji. Department of Agriculture, Fiji. Bulletin No. 35. 196 pp.
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