Pacific Island Ecosystems at Risk (PIER)

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Alpinia purpurata
(Vieill.) K.Schum., Zingiberaceae
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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Risk assessment results:  Evaluate, score: 4 (Go to the risk assessment)

Common name(s): [more details]

English: red ginger

Fijian: thevunga

Hawaiian: ‘awapuhi ‘ula‘ula

Kwara‘ae: fi‘i ange, forlata

Maori (Cook Islands): kāopi kura, kāopi muramura, kaopu‘i muramura, kaopui kutekute, kōpī muramura

Nangu: aluko

Niuean: kamapui futi kula, keuila

Rotuman: kapui riri‘i

Samoan: teuila, teuila, teuila, teuila mćmć, teuila mćmć

Tahitian: opuhi uteute

Tokelauan: teuila

Tongan: teuila, teuila pāpālangi, tevunga

Yapese: telan

Habit:  herb

Description:  "Leafy shoots 1-5 m tall (up to 7 m tall in cultivated forms).  Leaves oblong, 30-70 (-80) cm long, 10-22 cm wide, glabrous, apex short-acuminate, sessile, ligules unequally 2-lobed, (5-) 7-20 mm long, pubescent, sheaths pubescent at apex margins.  Inflorescences terminal on leafy shoots, unbranched (occasionally branched in cultivated forms), basically cylindrical, 15-30 cm long, usually elongating considerably with age, primary bracts usually red (occasionally pink or white in cultivated forms), ovate or broadly obovate, 2.5-3 cm long at anthesis, elongating to 4-6 cm long in fruit, apex mucronate, persistent, rachis glabrous, glabrate, or rarely pubescent, cincinni 1-2 (-5)-flowered, bracteoles reddish, tubular, 6-10 mm long, with a subapical spur; floral tube 2-2.7 mm long, calyx reddish, 1.7-2 (-2.7) cm long, glabrous, each lobe with a subapical spur; corolla white, exserted just beyond primary bracts; labellum white, with 2 distinct lateral lobes and a central segment that is distinctly 3-lobed or subentire, about as long as corolla lobes; stamen 6-7 mm long; ovary 3-4 mm long, glabrous.  Capsules nearly globose, 2-3 cm in diameter.  Seeds ca 3 mm long, angled"  (Wagner et al., 1999; p. 1618).

Habitat/ecology:  In Hawai‘i, "introduced as an ornamental as early as 1928, now naturalized" (Wagner et al., 1999; p. 1618).  In Fiji, "naturalized in abandoned garden areas, along trails and streams in forests, and on the edges of mangrove swamps, at elevations up to 500 m" (Smith, 1975; p. 211).

Propagation:  Seed

Native range:  New Caledonia, New Hebrides, Solomon Islands, Bismarck Archipelago and Bougainville. Widely cultivated and naturalized in the tropics (Wagner et al., 1999; p. 1618).

Presence:

Pacific
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
American Samoa
Manu‘a Islands
Ofu Island   Yuncker, T. G. (1945) (p. 30)
Voucher cited: Yuncker 9460 (BISH)
American Samoa
Manu‘a Islands
Ofu Island   Bishop Museum (Honolulu) (1939) (voucher ID: BISH 699993)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
American Samoa
Manu‘a Islands
Ta‘u Island   Yuncker, T. G. (1945) (p. 30)
Voucher cited: Yuncker 9177 (BISH)
American Samoa
Manu‘a Islands
Ta‘u Island   Bishop Museum (Honolulu) (1939) (voucher ID: BISH 699992)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
American Samoa
Tutuila Islands
Tutuila Island   Bishop Museum (Honolulu) (1975) (voucher ID: BISH 430371)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Rota Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Cook Islands
Southern Cook Islands
Aitutaki Atoll introduced
cultivated
Space, James C./Flynn, Tim (2002) (p. 73)
Cook Islands
Southern Cook Islands
‘Atiu Island introduced
cultivated
Space, James C./Flynn, Tim (2002) (p. 78)
Cook Islands
Southern Cook Islands
Mangaia Island cultivated
Sykes, Bill (year unknown)
Cook Islands
Southern Cook Islands
Rarotonga Island introduced
cultivated
Space, James C./Flynn, Tim (2002) (p. 63)
Ecuador (Galápagos Islands)
San Cristóbal Group
San Cristóbal Island introduced
cultivated
Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands)
Santa Cruz Group
Santa Cruz Island introduced
cultivated
Charles Darwin Foundation (2008)
Federated States of Micronesia
Chuuk Islands
Puluwat (Pwonowót) Atoll introduced
cultivated
Manner, H. I./Mallon, E. (1989) (p. 41)
Voucher cited: Manner & Mallon 200 (GUAM)
Federated States of Micronesia
Chuuk Islands
Weno (Moen) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Federated States of Micronesia
Kosrae Island
Kosrae Island introduced
cultivated
Lorence, David H./Flynn, Timothy (2010) (p. 11)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
cultivated
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 62)
Voucher cited: F.R. Fosberg 58448 (BISH)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   Bishop Museum (Honolulu) (1978) (voucher ID: BISH 438250)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Federated States of Micronesia
Yap Islands
Lamotrek Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Federated States of Micronesia
Yap Islands
Lamotrek Atoll   Bishop Museum (Honolulu) (1965) (voucher ID: BISH 595562)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Federated States of Micronesia
Yap Islands
Yap (Waqab) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Fiji
Fiji Islands
Kandavu Island introduced
invasive
cultivated
Smith, Albert C. (1979) (p. 211)
Voucher cited: Smith 8
Fiji
Fiji Islands
Kandavu Island   Bishop Museum (Honolulu) (1933) (voucher ID: BISH 699985)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Rotuma Island   Bishop Museum (Honolulu) (1938) (voucher ID: BISH 617418)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Rotuma Island   Bishop Museum (Honolulu) (1991) (voucher ID: BISH 637987)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Vanua Levu Island introduced
invasive
cultivated
Smith, Albert C. (1979) (p. 211)
Voucher cited: Smith 6827
Fiji
Fiji Islands
Vanua Levu Island   Bishop Museum (Honolulu) (1947) (voucher ID: BISH 699986)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Viti Levu Island introduced
invasive
cultivated
Smith, Albert C. (1979) (p. 211)
Vouchers cited: Gillespie 4384, DA 15449, DA 12154, DA 12336, DA 16230
Fiji
Fiji Islands
Viti Levu Island   Bishop Museum (Honolulu) (1967) (voucher ID: BISH 34045)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Viti Levu Island   Bishop Museum (Honolulu) (1953) (voucher ID: BISH 699987)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Viti Levu Island   Bishop Museum (Honolulu) (1924) (voucher ID: BISH 699988)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Viti Levu Island   Bishop Museum (Honolulu) (1927) (voucher ID: BISH 699989)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Fiji
Fiji Islands
Viti Levu Island   Bishop Museum (Honolulu) (1953) (voucher ID: BISH 699990)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
French Polynesia
Gambier Islands
Akamaru Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
cultivated
Lorence, David H./Wagner, Warren L. (2008)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
cultivated
Lorence, David H./Wagner, Warren L. (2008)
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
cultivated
Lorence, David H./Wagner, Warren L. (2008)
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
cultivated
Lorence, David H./Wagner, Warren L. (2008)
French Polynesia
Society Islands
Huahine Island introduced
Fosberg, F. R. (1997) (p. 154)
French Polynesia
Society Islands
Maupiti (Maurua) Island introduced
cultivated
Fosberg, F. R./Sachet, M.-H. (1987) (p. 23)
French Polynesia
Society Islands
Moorea Island introduced
Fosberg, F. R. (1997) (p. 154)
French Polynesia
Society Islands
Moorea Island introduced
cultivated
University of California (2006)
"Grown ornamentally for its bright, showy bracts".
French Polynesia
Society Islands
Moorea Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
cultivated
Welsh, S. L. (1998) (pp. 378-379)
Vouchers cited: Moore 115, Moore 385
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Society Islands
Taha‘a Island introduced
Fosberg, F. R. (1997) (p. 154)
French Polynesia
Society Islands
Taha‘a Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
French Polynesia
Society Islands
Tahiti Island introduced
cultivated
Welsh, S. L. (1998) (pp. 378-379)
Vouchers cited: BRY 24812, Florence 2585
French Polynesia
Society Islands
Tahiti Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Voucher cited: J. Florence 2585 (PAP)
Très commune à Tahiti où on cultive aussi une variété à bractées roses.
French Polynesia
Society Islands
Tahiti Island   Bishop Museum (Honolulu) (1982) (voucher ID: BISH 493479)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
cultivated
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2011)
Cultivée
Guam
Guam Island
Guam Island probably introduced
cultivated
Stone, Benjamin C. (1970) (p. 110)
Guam
Guam Island
Guam Island   Bishop Museum (Honolulu) (1966) (voucher ID: BISH 595566)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Guam
Guam Island
Guam Island   Bishop Museum (Honolulu) (1962) (voucher ID: BISH 699984)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
State of Hawaii
Hawaiian Islands
Hawai‘i (Big) Island introduced
Wester, Lyndon (1992) (p. 124)
State of Hawaii
Hawaiian Islands
Kaua‘i Island introduced
invasive
cultivated
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1618)
Vouchers cited: St. John 12798, 12767 (BISH)
State of Hawaii
Hawaiian Islands
Moloka‘i Island introduced
invasive
cultivated
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1618)
State of Hawaii
Hawaiian Islands
O‘ahu Island introduced
Wester, Lyndon (1992) (p. 124)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
cultivated
Whistler, W. A./Steele, O. (1999) (p. 105)
Marshall Islands
Ratak Chain
Majuro (Mãjro) Atoll introduced
cultivated
abo. intro.
Vander Velde, Nancy (2003) (p. 56)
Nauru
Nauru Island
Nauru Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Nauru
Nauru Island
Nauru Island introduced
cultivated
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (p. 75)
Voucher cited: Fosberg 59710 (US)
New Caledonia
New Caledonia Archipelago
Île Grande Terre native
MacKee, H. S. (1994) (p. 141)
Vouchers cited: MacKee 7902, Lauri 120 (NOU)
Niue
Niue
Niue Island introduced
cultivated
Sykes, W. R. (1970) (p. 270)
Voucher cited: CHR 169972
Niue
Niue
Niue Island introduced
cultivated
Yuncker, T. G. (1943) (p. 40)
Voucher cited: Yuncker 10086 (BISH)
Niue
Niue
Niue Island introduced
invasive
cultivated
Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 38)
Palau
Palau (main island group)
Angaur Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 88)
Palau
Palau (main island group)
Babeldaob Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 84)
Palau
Palau (main island group)
Koror Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1987) (p. 111)
Palau
Palau (main island group)
Koror Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 79)
Palau
Palau (main island group)
Koror Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 58)
Palau
Palau (main island group)
Koror Island   Beleu National Museum (Republic of Palau. Koror.) (1974) (voucher ID: BNM 10049)
Taxon name on voucher: Alpinia purpurata (Viell.) K. Schum.
Palau
Palau (main island group)
Koror Island   Beleu National Museum (Republic of Palau. Koror.) (1968) (voucher ID: BNM 121)
Taxon name on voucher: Alpinia purpurata (Viell.) K. Schum.
Palau
Palau (main island group)
Malakal Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 79)
Palau
Palau (main island group)
Ngerkebesang Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 79)
Papua New Guinea
Bismarck Archipelago
Bismarck Archipelago native
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1618)
Papua New Guinea
Bismarck Archipelago
Bismarck Archipelago   Peekel, P. G. [translated by E. E. Henty] (1984) (p. 103)
Papua New Guinea
Bougainville Islands
Bougainville Island   Foreman, D. B. (1971) (p. 31)
Samoa
Western Samoa Islands
Savai‘i Island introduced
invasive
Space, James C./Flynn, Tim (2002) (p. 69)
Samoa
Western Samoa Islands
Savai‘i Island   Bishop Museum (Honolulu) (1931) (voucher ID: BISH 699994)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Samoa
Western Samoa Islands
Upolu Island introduced
invasive
Space, James C./Flynn, Tim (2002) (p. 65)
Samoa
Western Samoa Islands
Upolu Island   Bishop Museum (Honolulu) (1979) (voucher ID: BISH 439389)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Samoa
Western Samoa Islands
Upolu Island   Bishop Museum (Honolulu) (1929) (voucher ID: BISH 699995)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Solomon Islands
Solomon Islands
Mikira (San Cristobal) Island   Bishop Museum (Honolulu) (1932) (voucher ID: BISH 127396)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Solomon Islands
Solomon Islands
Ndeni (Santa Cruz) Island   Bishop Museum (Honolulu) (1972) (voucher ID: BISH 79936)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Solomon Islands
Solomon Islands
Solomon Islands native
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1618)
Solomon Islands
Solomon Islands
Solomon Islands introduced
cultivated
Hancock, I. R./Henderson, C. P. (1988) (p. 45)
Tonga
Tongatapu Group
Tongatapu Island introduced
invasive
Space, James C./Flynn, Tim (2001) (p. 50)
Tonga
Tongatapu Group
Tongatapu Island   Bishop Museum (Honolulu) (1959) (voucher ID: BISH 41398)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Tonga
Tongatapu Group
Tongatapu Island   Bishop Museum (Honolulu) (1953) (voucher ID: BISH 127314)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Tonga
Vava‘u Group
Niuatoputapu Island   Bishop Museum (Honolulu) (1976) (voucher ID: BISH 456103)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.
Vanuatu
New Hebrides Islands
Vanuatu (Republic of) native
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1618)
Pacific Rim
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Singapore
Singapore
Singapore (Republic of) introduced
cultivated
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 12)
Cultivated only
Also reported from
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
United States (continental except west coast)
United States (other states)
USA (Florida)   Bishop Museum (Honolulu) (1903) (voucher ID: BISH 634305)
Taxon name on voucher: Alpinia purpurata (Vieill.) K.Schum.

Additional information:
Additional online information about Alpinia purpurata is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Alpinia purpurata as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Alpinia purpurata may be available from the Germplasm Resources Information Network (GRIN).

References:

Beleu National Museum (Republic of Palau. Koror.). 1968. Voucher specimen #BNM121(Blackburn 185).

Beleu National Museum (Republic of Palau. Koror.). 1974. Voucher specimen #BNM10049(Otobed, D.O. P-10049).

Bishop Museum (Honolulu). 1903. Voucher specimen #BISH 634305 (Seale, A.; collector's voucher number unknown).

Bishop Museum (Honolulu). 1924. Voucher specimen #BISH 699988 (Bryan Jr, E.H. 217).

Bishop Museum (Honolulu). 1927. Voucher specimen #BISH 699989 (Gillespie, J.W. 4384).

Bishop Museum (Honolulu). 1929. Voucher specimen #BISH 699995 (Christophersen, E. 346).

Bishop Museum (Honolulu). 1931. Voucher specimen #BISH 699994 (Christophersen, E. 2009).

Bishop Museum (Honolulu). 1932. Voucher specimen #BISH 127396 (Brass, L.J. 2570).

Bishop Museum (Honolulu). 1933. Voucher specimen #BISH 699985 (Smith, A.C. 8).

Bishop Museum (Honolulu). 1938. Voucher specimen #BISH 617418 (St.John, H. 19775).

Bishop Museum (Honolulu). 1939. Voucher specimen #BISH 699992 (Yuncker, T.G. 9177).

Bishop Museum (Honolulu). 1939. Voucher specimen #BISH 699993 (Yuncker, T.G. 9460).

Bishop Museum (Honolulu). 1947. Voucher specimen #BISH 699986 (Smith, A.C. 6827).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 127314 (Yuncker, T.G. 15263).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 699987 (Smith, A.C. 7216).

Bishop Museum (Honolulu). 1953. Voucher specimen #BISH 699990 (Smith, A.C. 7216).

Bishop Museum (Honolulu). 1959. Voucher specimen #BISH 41398 (Soakai, E. 749).

Bishop Museum (Honolulu). 1962. Voucher specimen #BISH 699984 (Stone, B.C.M. 4119).

Bishop Museum (Honolulu). 1965. Voucher specimen #BISH 595562 (Fosberg, F.R. 46815).

Bishop Museum (Honolulu). 1966. Voucher specimen #BISH 595566 (Evans, M. 1602).

Bishop Museum (Honolulu). 1967. Voucher specimen #BISH 34045 (Kuruvoli, I.T. 15449).

Bishop Museum (Honolulu). 1972. Voucher specimen #BISH 79936 (Powell, J.M. 19845).

Bishop Museum (Honolulu). 1975. Voucher specimen #BISH 430371 (Whistler, W.A. W2877).

Bishop Museum (Honolulu). 1976. Voucher specimen #BISH 456103 (Kirch, P.V. 325).

Bishop Museum (Honolulu). 1978. Voucher specimen #BISH 438250 (Fosberg, F.R. 58448).

Bishop Museum (Honolulu). 1979. Voucher specimen #BISH 439389 (Cox, P.A. 163).

Bishop Museum (Honolulu). 1982. Voucher specimen #BISH 493479 (Florence, J. 2585).

Bishop Museum (Honolulu). 1991. Voucher specimen #BISH 637987 (McClatchey, W.C. 112).

Charles Darwin Foundation. 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Charles Darwin Foundation, Galapagos, Ecuador.

Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species. Raffles Museum of Biodiversity Research, National University of Singapore. 273 pp.

Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. 2011. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP). (online resource).

Foreman, D. B. 1971. A check list of the vascular plants of Bougainville, with descriptions of some common forest trees. Botany Bulletin No. 5. Division of Botany, Department of Forests. Lae, New Guinea. 194 pp.

Fosberg, F. R. 1997. Preliminary checklist of the flowering plants and ferns of the Society Islands. Ed. by David R. Stoddart. U. Cal. Berkeley.

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This page was created on 1 JAN 1999 and was last updated on 12 OCT 2006.