Pacific Island Ecosystems at Risk (PIER)

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Achyranthes aspera
L., Amaranthaceae
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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Common name(s): [more details]

Austral: erofai

Chamorro: chichiton, chichitun, lasocata, lasogado

Chinese: tu niu xi

Chuukese: ikkakeeke, likakeeke, likkakééké, liko, niké, niko, niroke, seetan, uoko, uökö

English: achyranthes, burweed, chaff-flower, chaffbur, crocus stuff, crokars staff, devil's horsewhip, prickly chaff-flower

Fijian: sono ivi

French: herbe à Bengalis, herbe sergen, queue de rat

Kapingamarangi: tikeng tutui

Mangarevan: aerofai, pukawapuataratara

Maori (Cook Islands): kōpū varu, piripiri, piripiri, tama ‘enua, tumu ‘enua

Marquesan: moto ouaua, otunatuna, teita‘a

Marshallese: kaleklek, kaleklek, kaloklok

Mokilese: suga dugokok

Niuean: lamoa fisi, talamoa, talamoa fisi, talamoa fiti

Palauan: louch beluu, louch el beluu

Pohnpeian: osen likendinikep

Samoan: la‘au talatala, la‘au tālātālā, lau tamatama, lau tamatama, lau tamatama, tamatama, tamatama

Satawalese: eeg‘gohur

Spanish: cadillo chichoborugo, cadillo de mazorca, mazotillo, rabo de chancho, rabo de ratón

Tahitian: aero fai, aerofai, aerofai, aerofai, aerofai, airovai, aruhai, erofai

Tokelauan: lau tamatama

Tongan: tamatama

Tuamotuan: aero fai, arofai

Tuvaluan: sisi vao, tamatama

Ulithian: koi

Wallisian: tamatama

Woleaian: gugu

Yapese: gogo

Habit:  herb

Description: 

Genus: "Erect or ascending herbs or shrubs; leaves opposite, the blades entire; inflorescences terminal and axillary, spicate, erect, many-flowered; becoming elongate, with only a few flowers open at the same time; flowers hermaphrodite, solitary in axils of acute, membranous, persistent bracts, subtended by 2 bracteoles, deflexed after anthesis, the bracteoles consisting of a long spine and bearing on each side of base a shorter, membranous, nerveless wing; tepals 5, spreading during anthesis, before and after anthesis erect, membranous or herbaceous, 1- or more-nerved, acute, sometimes pungent in fruit; stamens 5, much shorter than perianth, the filaments proximally connate into a short cup, the free parts alternating with short, broad pseudostaminodes, the anthers oblong, 2-celled (4-loculed); ovary glabrous, the ovule 1, pendent from a long funicle, the style filiform, short, the stigma capitate, utricle falling off together with perianth and bracteoles, ellipsoid, indehiscent, 1-seeded, with truncate or depressed apex, thin-walled, the seed erect."  (Smith, 1981; p. 290)

Species: "A coarse, ligneous herb or shrub 0.8-4 m high, sometimes almost treelike. The tepals are white to pale lavender, the filaments while to rich pink, and the fruit orange to reddish purple or brown."  (Smith, 1981; p. 290).

"A coarse, rambling or erect widely branched annual or short-lived perennial with a semi-woody base; taproot long with lateral branches throughout; stem 0.5 to 2 m tall, angularly-ribbed, generally square, more or less densely hairy and thickened above the nodes; leaves opposite, oblong-obovate to elliptic or obovate from an acute or obtuse base; tip acuminate, acute, obtuse or rounded; blade entire, flat or somewhat wavy, more or less pubescent, 2 to 10 cm long, 0.7 to 5 cm wide; petiole 0.5 to 1.5 cm long; inflorescence terminal spikes, rigid, 10 to 50 cm long excluding the peduncle, with paired branches below; rachis robust, angularly-ribbed, more or less hairy; flowers small, green, perfect, densely arranged at top of spike, less clustered in the center, scattered and often in pairs near the base; subtended by long-acuminate bracts or bracteoles, 2 to 3.5 mm long, stiff and spiny, erect before anthesis reflexed later, persistent; sepals 5, green with pale margins, ovate-lanceolate, acuminate, 3.5 to 5.5 mm long during anthesis, five filaments 2.2 to 3.5 mm long; pseudo-staminodia with or without dorsal, truncate, long-fringed scales; ovary top-shaped with 1- to 2-mm style; fruit and utricle (a small prickly bur), 2.5 to 2.8 mm long, rounded at the base; enclosed by persistent perianth and bracts, 4 to 5 mm long, detaching easily from rachis; seed 2 to 3 mm long, 1 to 1.5 mm wide, truncate above, reddish to dark brown and shiny, enclosed in chaffy calyx parts that remain attached.  This species is readily distinguished by the opposite leaves, branched stem and spiny bracts that are erect before flowering but then become reflexed and readily adhere to animals and clothing"  (Holm et al., 1997; p. 13).

Description from the Flora of China online.
Description from the Flora of North America onkline.

Habitat/ecology:  "It frequently occurs in waste areas, and along roadsides, foot paths, railroads and sand dunes.  It often infest fence rows, open woodland, and the boarders of forests and coffee fields.  It has adapted to a wide range of environments (Holm et al., 1997; p. 13).

In Fiji, "occurs from near sea level to about 900 m elevation as an abundantly naturalized weed on rocky shores, limestone islets, and grassy slopes, in coastal thickets, cultivated areas, and along roadsides and forest trails" (Smith, 1981; p. 290).  In Hawai‘i, "naturalized in low elevation, open, dry disturbed habitats, such as along roadsides"  (Wagner et al., 1999; pp. 180-181).  In Niue, "locally fairly common in waste places, especially near tracks" (Sykes, 1970; p. 40). In Guam, "a troublesome, frequent weed of waste ground, trails, etc." [probably native] (Stone, 1970; p. 270). In Papua New Guinea, "a weed of pastures, roadsides and waste places; also in natural situations, behind beaches, in savannah grassland. Widely spread, at low elevations in areas of seasonal climate" (Henty & Pritchard, 1975; p. 51).  In Tonga, "a common plantation and waste area weed" (Yuncker, 1959; p. 108).  In Papua New Guinea, "a weed of dry open places, such as roadsides and gardens, especially in areas subject to a dry season, at 0-200(?) altitude"  (Womersley, 1978; p. 15).  In Tonga, "a common plantation and waste area weed" (Yuncker, 1959; p. 108).

Propagation:  Seed

Native range:  "Probably indigenous in southeastern Asia and now a ubiquitous weed, found in Pacific archipelagoes eastward to the Tuamotus" (Smith, 1981; p. 290).

"Native or aboriginally introduced to Polynesia" (Whistler, 1988; p. 6). Fosberg et al. (1979; p. 67) lists as native to Micronesia.

Presence:

Pacific
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
American Samoa
Manu‘a Islands
Ofu Island   Whistler, W. A. (1980) (p. 159)
American Samoa
Manu‘a Islands
Olosega Island   Whistler, W. A. (1980) (p. 159)
American Samoa
Manu‘a Islands
Olosega Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2001) (voucher ID: PTBG 35216)
Taxon name on voucher: Achyranthes aspera L.
American Samoa
Manu‘a Islands
Ta‘u Island   Whistler, W. A. (1980) (p. 159)
American Samoa
Manu‘a Islands
Ta‘u Island   Yuncker, T. G. (1945) (p. 37)
Vouchers cited: Garber 563 (BISH), Garber 568 (BISH), W. Harris 322 (BISH)
American Samoa
Manu‘a Islands
Ta‘u Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1990) (voucher ID: PTBG 9104)
Taxon name on voucher: Achyranthes aspera
American Samoa
Tutuila Islands
Aunu‘u Island   Whistler, W. A. (1980) (p. 159)
American Samoa
Tutuila Islands
Tutuila Island   Whistler, W. A. (1980) (p. 159)
American Samoa
Tutuila Islands
Tutuila Island introduced
invasive
Setchell, William Albert (1924) (p. 96)
Australia (Pacific offshore islands)
Lord Howe Island
Lord Howe Island native
Orchard, Anthony E., ed. (1994) (p. 18)
"Probably native on coastal cliffs and similar habitats but becoming invasive and weedy on disturbed soil". Vouchers cited: A.N. Rodd 1452 (K, NSW), J.C. Game 69/274e (K), P.S. Green 1650 (A, K)
Australia (Pacific offshore islands)
Norfolk Islands
Norfolk Island native
Orchard, Anthony E., ed. (1994) (p. 7)
"Probably native on coastal cliffs and similar habitats but becoming invasive and weedy on disturbed soil". Vouchers cited: O. Evans (K), P.S. Green 1446 (A), J.H. Maiden & J.L. Boorman (K, NSW)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Agrigan (Agrihan) Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Agrigan (Agrihan) Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 23)
Voucher cited: Fosberg 31596 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Aguijan (Agiguan, Aguihan) Island introduced
Raulerson, L. (2006) (p. 28)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Asuncion Island native
Falanruw, M. V. C. (1989) (p. 8)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Asuncion Island introduced
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1977) (p. 29)
Voucher cited: Falanruw 3124 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Guguan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1977) (p. 29)
var. fruticosa (Lam.) Boerl. Voucher cited: Falanruw 3102 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Pagan Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Pagan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 23)
Voucher cited: Moore 338 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Rota Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Rota Island native
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane (2000) (p. Voucher specimen)
Voucher: BMW 5812 (BISH, BRI, GUAM, CANB)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Saipan Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Tinian Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Cook Islands
Southern Cook Islands
Aitutaki Atoll introduced
invasive
Florence, J. (2004) (pp. 44-47)
Cook Islands
Southern Cook Islands
‘Atiu Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
Cook Islands
Southern Cook Islands
Mangaia Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Ma‘uke Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Miti‘aro Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Rarotonga Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
Ecuador (Galápagos Islands)
Isabela Group
Isabela Island introduced
invasive
Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands)
Isabela Group
Volcán Sierra Negra, Isabela Island introduced
invasive
Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands)
San Cristóbal Group
San Cristóbal Island introduced
invasive
Charles Darwin Foundation (2008)
Ecuador (Galápagos Islands)
Santa Cruz Group
Santa Cruz Island introduced
invasive
Charles Darwin Foundation (2008)
Federated States of Micronesia
Chuuk Islands
Lukunor (Núkúnooch) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Murilo (Mwiriné) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Nama Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Namoluk (Nómwunuuk) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Namoluk (Nómwunuuk) Atoll   Marshall, Mae/Fosberg, F. R. (1975) (p. 33)
Voucher cited: Marshall 76 (US)
Federated States of Micronesia
Chuuk Islands
Namonuito (Namonweito, Nómwun Wiité) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Nomwin (Nómwiin) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Pis Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Chuuk Islands
Puluwat (Pwonowót) Atoll native
Manner, H. I./Mallon, E. (1989) (p. 44)
Voucher cited: Manner & Mallon 145 (GUAM)
Federated States of Micronesia
Chuuk Islands
Satawan Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Kosrae Island
Kosrae Island introduced
invasive
Josekutty, P. C./Wakuk, E. E./Joseph, M. J. (2002) (p. 62)
Federated States of Micronesia
Pohnpei Islands
Kapingamarangi Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Pohnpei Islands
Mokil (Mwokil) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
invasive
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 66)
Vouchers cited: A. Dores 202 (NY, PTBG), F.R. Fosberg 26061 (BISH), W.A. Niering 546 (BISH), S. Perlman 21412 (PTBG)
Naturalized
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 43549)
Taxon name on voucher: Achyranthes aspera var. aspera
Federated States of Micronesia
Yap Islands
Eauripik Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Yap Islands
Fais Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Yap Islands
Fais Island introduced
invasive
Fosberg, F. R./Evans, M. (1969) (p. 6)
Voucher cited: Fosberg 46711 (US)
Federated States of Micronesia
Yap Islands
Faraulep (Faraulap) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Yap Islands
Ifalik (Ifaluk) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Yap Islands
Satawal Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Yap Islands
Satawal Island introduced
invasive
Fosberg, F. R. (1969) (p. 7)
Voucher cited: Fosberg 46914 (US)
Federated States of Micronesia
Yap Islands
Sorol Atoll   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Federated States of Micronesia
Yap Islands
Ulithi Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Federated States of Micronesia
Yap Islands
Woleai Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Fiji
Fiji Islands
Bacon Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Bryan 538
Fiji
Fiji Islands
Kandavu Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Tothill 665
Fiji
Fiji Islands
Koro Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Tothill 664
Fiji
Fiji Islands
Mango Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Bryan 563
Fiji
Fiji Islands
Matuku Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Bryan 234
Fiji
Fiji Islands
Mbengga Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Weiner 223
Fiji
Fiji Islands
Moala Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Smith 1396
Fiji
Fiji Islands
Taveuni Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Smith 8114
Fiji
Fiji Islands
Vanua Levu Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: DA 16055
Fiji
Fiji Islands
Vanua Mbalavu Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Voucher cited: Tothill 666
Fiji
Fiji Islands
Viti Levu Island introduced
invasive
Smith, Albert C. (1981) (p. 290)
Vouchers cited: DA 11755, DA 9683, Greenwood 61, DA 13974, DA 11866, Greenwood 61A, DA 9332, H.B.R. Parham 43
French Polynesia
Gambier Islands
Agakauitai Island introduced
St. John, Harold (1988) (p. 13)
Observed
French Polynesia
Gambier Islands
Aukena Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Gambier Islands
Aukena Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Gambier Islands
Aukena Island introduced
St. John, Harold (1988) (p. 13)
Voucher cited: St. John 14662 (BISH)
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Gambier Islands
Mangareva (Gambier) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: S.P. Perlman & K.R. Wood 15001 (PAP)
Adventice
French Polynesia
Marquesas Islands
Fatu Hiva Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 22468)
Taxon name on voucher: Achyranthes aspera var. aspera L.
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Mohotani (Mohotane, Mohotane) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Mohotani (Mohotane, Mohotane) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Mohotani (Mohotane, Mohotane) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1964) (voucher ID: PTBG 8938)
Taxon name on voucher: Achyranthes aspera var. aspera
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Bora Bora Island introduced
invasive
Welsh, S. L. (1998) (p. 25)
Voucher cited: Grant 4873
French Polynesia
Society Islands
Bora Bora Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Society Islands
Bora Bora Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 12015 (PAP)
Adventice
French Polynesia
Society Islands
Huahine Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Society Islands
Huahine Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence & R. Tahuaitu 11556 (PAP); var. velutina (W.J. Hooker & Arnott) C. Townsend: F.R. Fosberg 61174 (PAP), D. Baré 6 (PAP)
Adventice
French Polynesia
Society Islands
Maiao (Maiaoiti, Mauiti, Tupuaemanu) Island native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Society Islands
Maupiti (Maurua) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. aspera. Voucher cited: J.-F. Butaud & F.A. Jacq 41 (PAP)
Adventice
French Polynesia
Society Islands
Mehetia (Mahetia, Meetia) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Society Islands
Moorea Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Welsh, S. L. (1998) (p. 25)
Vouchers cited: Moore 346, BRY 26422
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Taha‘a Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Welsh, S. L. (1998) (p. 25)
Vouchers cited: Setchell & Parks 320, Grant 4006, Grant 4676
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Society Islands
Tetiaroa Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 3269 (PAP)
French Polynesia
Society Islands
Tupai Island native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: M.-H. Sachet 1996 (PAP)
French Polynesia
Tuamotu Archipelago
Ana‘a Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Apataki (Apa‘ataki) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Hao (Hau) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Hiti Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Katiu (Katui) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Kauehi (Kawehi) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Kaukura Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Makatea (Ma‘atea) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Tuamotu Archipelago
Makatea (Ma‘atea) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 3025 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Marutea Sud (Marutea South) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Matureivavao (Maturei Vavao) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Napuka Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Niau Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Nukutavake Island native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Tuamotu Archipelago
Paraoa Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Rangiroa (Rairoa) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Vouchers cited: J. Florence 9972 (PAP), J. Florence 9954 (PAP), J. Florence 10021 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Raro‘ia Atoll native
Meyer, Jean-Yves/Fourdrigniez, Marie (2008) (p. 10)
Var. velutina
French Polynesia
Tuamotu Archipelago
Raro‘ia Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Reao Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Taenga Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Taiaro Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Takapoto Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: M.-H. Sachet 2018 (PAP)
French Polynesia
Tuamotu Archipelago
Takume (Takurea) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Tenararo (Tenaroa) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
French Polynesia
Tuamotu Archipelago
Tenarunga Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 3269 (PAP)
French Polynesia
Tuamotu Archipelago
Tepoto Nord Island native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 3269 (PAP)
French Polynesia
Tuamotu Archipelago
Tikehau (Tikahau) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Vouchers cited: J. Florence 7070 (PAP), J. Florence 7124 (PAP)
French Polynesia
Tuamotu Archipelago
Tikei (Tikai, Tiku) Island native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 3269 (PAP)
French Polynesia
Tuamotu Archipelago
Toau (Tovau) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 3269 (PAP)
French Polynesia
Tuamotu Archipelago
Tuanake Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 3269 (PAP)
French Polynesia
Tuamotu Archipelago
Tureia Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: M.-H. Sachet 1996 (PAP)
French Polynesia
Tuamotu Archipelago
Vahanga Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: M.-H. Sachet 1996 (PAP)
French Polynesia
Tuamotu Archipelago
Vanavana (Vana Vana) Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: M.-H. Sachet 1996 (PAP)
French Polynesia
Austral (Tubuai) Islands
Maria Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence 10171 (PAP)
French Polynesia
Austral (Tubuai) Islands
Raivavae (Raevavae) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Austral (Tubuai) Islands
Raivavae (Raevavae) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. aspera. Vouchers cited: J. Florence 5692 (PAP), J. Florence 5804 (PAP), J. Florence 5864 (PAP), J. Florence & W.R. Sykes 11148 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rapa Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Austral (Tubuai) Islands
Rapa Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 6319 (PAP), J. Florence 6331 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rapa Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 42025)
Taxon name on voucher: Achyranthes aspera
French Polynesia
Austral (Tubuai) Islands
Rapa Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2002) (voucher ID: PTBG 42304)
Taxon name on voucher: Achyranthes aspera
French Polynesia
Austral (Tubuai) Islands
Rimatara (Rimitara) Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Austral (Tubuai) Islands
Rimatara (Rimitara) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 5568 (PAP), J. Florence 5592 (PAP), J. Florence 5973 (PAP), J. Florence 6150 (PAP), J. Florence 9198 (PAP), J. Florence 9207 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Tubuai Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
French Polynesia
Austral (Tubuai) Islands
Tubuai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
Guam
Guam Island
Guam Island native
Stone, Benjamin C. (1970) (p. 270)
Vouchers cited: Stone 4758 (GUAM), Stone 5269 (GUAM)
Guam
Guam Island
Guam Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Guam
Guam Island
Guam Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1994) (voucher ID: PTBG 22362)
Taxon name on voucher: Achyranthes aspera var. aspera
Guam
Guam Island
Guam Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1954) (voucher ID: PTBG 26636)
Taxon name on voucher: Achyranthes aspera var. aspera
State of Hawaii
Hawaiian Islands
Hawai‘i (Big) Island introduced
invasive
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 180-181)
State of Hawaii
Hawaiian Islands
Lāna‘i Island introduced
Wester, Lyndon (1992) (p. 125)
State of Hawaii
Hawaiian Islands
Moloka‘i Island introduced
invasive
Staples, George W./Imada, Clyde T./Herbst, Derral R. (2002) (p. 4)
Voucher cited: N. Matayoshi s.n. (BISH 652141)
State of Hawaii
Hawaiian Islands
O‘ahu Island introduced
invasive
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (pp. 180-181)
Japan (offshore islands)
Ogasawara (Bonin) Islands
Bonin (Ogasawara) Islands introduced
Kato, Hidetoshi (2007)
Var. aspera
Kiribati
Line Islands
Tabuaeran (Fanning) Atoll introduced
Wester, Lyndon (1985) (p. 21)
Collected by Arundel but not recorded since.
Kiribati
Tungaru (Gilbert) Islands
Tarawa Atoll   Swarbrick, John T. (1997) (p. 88)
Presence not verified
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll introduced
invasive
St. John, Harold (1960) (p. 325)
Voucher cited: Fosberg 24348 (US)
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 9)
Voucher cited: Fosberg 24348 (US)
New Caledonia
Îles Bélep (Belep Islands)
Île Art introduced
invasive
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: MacKee 19449
New Caledonia
Îles Chesterfield
Îles Chesterfield introduced
invasive
abo. intro.
MacKee, H. S. (1994) (pp. 10-11)
Vouchers cited: Anon s.n., Kusser in MacKee 33998, Kusser in MacKee 34009, Kusser in MacKee 34019
New Caledonia
Récifs D'Entrecasteaux (D'Entrecasteaux Reefs)
Île Huon introduced
invasive
abo. intro.
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: Jacquier in MacKee 44249
New Caledonia
Récifs D'Entrecasteaux (D'Entrecasteaux Reefs)
Île Surprise introduced
invasive
abo. intro.
MacKee, H. S. (1994) (pp. 10-11)
Vouchers cited: Rolland in MacKee 15953, Boulet in MacKee 31459
New Caledonia
Hunter Island
Île Hunter introduced
invasive
abo. intro.
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: Monzier in Veillon 6923
New Caledonia
Îles Loyauté (Loyalte Islands)
Île Maré introduced
invasive
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: Sarasin 488 (Z)
New Caledonia
Îles Loyauté (Loyalte Islands)
Île Walpole introduced
invasive
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: Hmakone & Sam in MacKee 25850
New Caledonia
Îles Loyauté (Loyalte Islands)
Île Walpole native
Renevier, Alain/Cherrier, Jean-Francois (1991) (p. 15)
Voucher cited: Hmakone & Sam in MacKee 25850
New Caledonia
New Caledonia
New Caledonia Islands probably introduced
abo. intro.
Gargominy, Oliver/Bouchet, Philipe/Pascal, Michel/Jaffre, Tanguy/Tourneu, Jean-Christophe (1996) (p. 379)
Indigenous or early introduction.
Espèce pantropicale peut-être indigène ou d'introduction ancienne.
New Caledonia
New Caledonia Archipelago
Île Améré introduced
invasive
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: MacKee 37528
New Caledonia
New Caledonia Archipelago
Île Grande Terre introduced
invasive
Tassin, Jacques (2005)
New Caledonia
New Caledonia Archipelago
Île Grande Terre introduced
invasive
abo. intro.
MacKee, H. S. (1994) (pp. 10-11)
Vouchers cited: Vieillard 1066, Delacour s.n., MacKee 12652, MacKee 31742,
New Caledonia
New Caledonia Archipelago
Île des Pins (Isle of Pines) introduced
invasive
Tassin, Jacques (2005)
New Caledonia
New Caledonia Archipelago
Île des Pins (Isle of Pines) introduced
invasive
abo. intro.
MacKee, H. S. (1994) (pp. 10-11)
Voucher cited: MacKee 13122
Niue
Niue
Niue Island   Sykes, W. R. (1970) (p. 40)
Vouchers cited: CHR 169889, CHR 169890
Niue
Niue
Niue Island probably native
abo. intro.
Whistler, W. A. (1988) (p. 6)
Indigenous or a Polynesian Introduction. Voucher cited: Whistler 10764
Niue
Niue
Niue Island introduced
invasive
abo. intro.
Whistler, W. A./Atherton, J. (1997) (p. 24)
Indigenous or a Polynesian Introduction. Voucher cited: Whistler 10764
Uncommon to occasional.
Niue
Niue
Niue Island introduced
invasive
Yuncker, T. G. (1943) (p. 51)
Vouchers cited: Yuncker 9630 (BISH), Yuncker 10111 (BISH)
"A common weedly plant in plantations and waste ares".
Niue
Niue
Niue Island introduced
invasive
abo. intro.
Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 49)
Indigenous or a Polynesian Introduction.
Palau
Hatohobei Group
Tobi Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island native
Fosberg, F. R./Canfield, Joan E. (1980) (p. 195)
Var. pubescens. Voucher cited: Canfield 231 (US)
Palau
Palau (Belau ) (main island group)
Babeldaob Island native
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 100)
Palau
Palau (Belau ) (main island group)
Kayangel (Ngcheangel) Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 8)
Voucher: Lorence 9803 (PTBG, BNM, NY, TAMU, US)
Palau
Palau (Belau ) (main island group)
Kayangel (Ngcheangel) Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2008) (voucher ID: PTBG 96)
Taxon name on voucher: Achyranthes aspera var. aspera
Palau
Palau (Belau ) (main island group)
Peleliu (Beliliou) Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Palau
Rock Islands (Chelbacheb)
Ngeanges Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Palau
Sonsorol Group
Merir Island native
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109)
Palau
Sonsorol Group
Sonsorol Island native
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 67)
Palau
Sonsorol Group
Sonsorol Island native
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 109)
Palau
Sonsorol Group
Sonsorol Island   Beleu National Museum (Republic of Palau. Koror.) (1968) (voucher ID: BNM 573)
Taxon name on voucher: Achyranthes aspera L.
Palau
Sonsorol Group
Sonsorol Island   Beleu National Museum (Republic of Palau. Koror.) (1968) (voucher ID: BNM 572)
Taxon name on voucher: Achyranthes aspera L.
Papua New Guinea
Bismarck Archipelago
Bismarck Archipelago invasive
Peekel, P. G. [translated by E. E. Henty] (1984) (p. 170)
Tropical weed; at forest edges and along bush tracks.
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island) invasive
Henty, E. E./Pritchard, G. H. (1975) (p. 51)
In New Guinea widely spread, at low elevations in areas of seasonal climate.
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island)   Waterhouse, D. F. (1997) (p. 59)
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island) probably introduced
invasive
Womersley, John S., ed. (1978) (pp. 14-15)
Philippines
Philippine Islands
Philippine Islands   Waterhouse, D. F. (1993) (p. 63)
Philippines
Philippine Islands
Philippine Islands probably introduced
invasive
Merrill, Elmer D. (1923) (p. 131)
At low and medium altitudes, a weed in open waste places; a pantropic weed, probably itroduced.
Pitcairn Islands
Pitcairn Islands
Henderson Island introduced
Fosberg, F. R./Sachet, M.-H./Stoddart, D. R. (1983) (p. 30)
Pitcairn Islands
Pitcairn Islands
Henderson Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. pubescens (C.H.B.A. Moquin-Tandon) C.C. Townsend. Vouchers cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10928 (PAP), J. Florence (legit M.L. Brooke) 11053 (PAP)
Adventice
Pitcairn Islands
Pitcairn Islands
Oeno Atoll native
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
var. velutina (W.J. Hooker & Arnott) C. Townsend. Voucher cited: J. Florence, A.J. Chepstow-Lusty & S. Waldren 10932 (PAP)
Pitcairn Islands
Pitcairn Islands
Pitcairn Island introduced
St. John, H. (1987) (p. 31)
Vouchers cited: Fosberg 11341, Fosberg & B. Christian 11236, Fosberg & R. Clark 11316, St. John 15042
Pitcairn Islands
Pitcairn Islands
Pitcairn Island introduced
invasive
Florence, J. (2004) (pp. 44-47)
Pitcairn Islands
Pitcairn Islands
Pitcairn Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 11020 (PAP)
Adventice
Samoa
Western Samoa Islands
Western Samoa Islands   Whistler, W. A. (1988) (p. 6)
Solomon Islands
Solomon Islands
Solomon Islands   Swarbrick, John T. (1997) (p. 88)
Solomon Islands
Solomon Islands
Solomon Islands native
Hancock, I. R./Henderson, C. P. (1988) (p. 43)
Tonga
Ha‘apai Group
Ha‘apai Islands introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Yuncker 15800, Soakai 301, Soakai 680, Whistler 6595
Weedy
Tonga
Ha‘apai Group
Lifuka Island   Yuncker, T. G. (1959) (p. 108)
Voucher cited: Yuncker 15800
Tonga
Ha‘apai Group
Tofua Island   Whistler, W. A. (1992)
Voucher cited: Buelow 538
Tonga
Tonga Islands
Tonga Islands   Yuncker, T. G. (1959) (p. 108)
"Common throughout the group".
Tonga
Tongatapu Group
‘Eua Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Lister, Parks 16380, Yuncker 15517, Hotta 5424, Sykes 109, Buelow 1855, Whistler 7159, Whistler 7274
Weedy
Tonga
Tongatapu Group
‘Eua Island   Yuncker, T. G. (1959) (p. 108)
Voucher cited: Yuncker 15517
Tonga
Tongatapu Group
Tongatapu Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Mosley s.n. (n.s.), Setchell & Parks 15373, Hurlimann 121, Yuncker 15137, Hotta 4257, Hotta 4262, Alexander 5833, Lawrence 140, Lawrence 223, Soakai 184, Soakai 495, Buelow 74, Whistler 6744, Whistler 6759
Weedy
Tonga
Tongatapu Group
Tongatapu Island   Yuncker, T. G. (1959) (p. 108)
Voucher cited: Yuncker 15137
Tonga
Tongatapu Group
Tongatapu Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1989) (voucher ID: PTBG 3291)
Taxon name on voucher: Achyranthes aspera
Tonga
Tongatapu Group
Tongatapu Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1989) (voucher ID: PTBG 3315)
Taxon name on voucher: Achyranthes aspera
Tonga
Vava‘u Group
Late Island native
Sykes, W. R. (1981) (p. 348)
Tonga
Vava‘u Group
Late Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Hotta 4767, Buelow & Sykes 234, Buelow & Sykes 267 (as A. canescens)
Weedy
Tonga
Vava‘u Group
Niuafo‘ou Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Voucher cited: Buelow 2058
Weedy
Tonga
Vava‘u Group
Niuatoputapu Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Kirch 206, Kirch 299, Whistler 6186
Weedy
Tonga
Vava‘u Group
Tafahi Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Voucher cited: Hurlimann 540
Weedy
Tonga
Vava‘u Group
Tafahi Island   Yuncker, T. G. (1959) (p. 108)
Voucher cited: Hürlimann 540
Tonga
Vava‘u Group
Vava‘u Island introduced
invasive
abo. intro.
Whistler, W. A. (1992) (p. 2)
Vouchers cited: Barclay s.n. (n.s.), Crosby 237, Hotta 4988, Dosksi 688
Weedy
Tonga
Vava‘u Group
Vava‘u Island   Yuncker, T. G. (1959) (p. 108)
Cited: Barclay.
Tonga
Vava‘u Group
Vava‘u Island   Burkill, I. H. (1900) (p. 51)
Tuvalu
Tuvalu (Ellis) Islands
Nui Atoll   Woodroffe, C. D. (1985) (p. 11)
Voucher cited: Woodroffe 126 (CHR)
Tuvalu
Tuvalu (Ellis) Islands
Tuvalu Islands   Swarbrick, John T. (1997) (p. 88)
Vanuatu
New Hebrides Islands
Vanuatu (Republic of)   Swarbrick, John T. (1997) (p. 88)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis and Futuna Islands   Whistler, W. A. (1988) (p. 6)
Pacific Rim
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Australia
Australia (continental)
Australia (continental) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Western Australia
Australia
Australia (continental)
New South Wales native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Australia
Australia (continental)
Northern Territory native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Australia
Australia (continental)
Queensland native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Cambodia
Cambodia
Cambodia (Kingdom of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Central America
Central America (Pacific rim)
Costa Rica (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Central America
Central America (Pacific rim)
El Salvador (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Central America
Central America (Pacific rim)
Guatemala (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Central America
Central America (Pacific rim)
Honduras (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Central America
Central America (Pacific rim)
Nicaragua (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
China
China
China (People's Republic of) native
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2013)
"Hillsides, waste places, roadsides, riverbanks; 800-2300 m".
China
China
Hong Kong native
Wu, Te-lin (2001) (p. 77)
Indonesia
Indonesia
Indonesia (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Malaysia
Malaysia
Malaysia (country of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Mexico
Mexico
Mexico (United Mexican States) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Chiapas, Quintana Roo
New Zealand
New Zealand
New Zealand (country) introduced
invasive
Webb, C. J./Sykes, W. R./Garnock-Jones, P. J. (1988) (p. 100)
"A garden weed at Silverdale".
Singapore
Singapore
Singapore (Republic of) uncertain if introduced
invasive
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 9)
Weed of uncertain origin
South America (Pacific rim)
South America (Pacific rim)
Colombia (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
South America (Pacific rim)
South America (Pacific rim)
Perú (Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Taiwan
Taiwan Island
Taiwan Island native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Thailand
Thailand
Thailand (Kingdom of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Vietnam
Vietnam
Vietnam (Socialist Republic of) native
U.S. Dept. Agr., Agr. Res. Serv. (2013)
Indian Ocean
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Australia (Indian Ocean offshore islands)
Christmas Island Group
Christmas Island native
Orchard, Anthony E., ed. (1993) (p. 16)
Australia (Indian Ocean offshore islands)
Cocos (Keeling) Islands
Cocos (Keeling) Islands native
Orchard, Anthony E., ed. (1993) (pp. 36, 38)
On both main atoll and North Keeling Island.
British Indian Ocean Territory
Chagos Archipelago
Diego Garcia Island   Stoddart, D. R./Taylor, J. D., eds. (1971) (p. 149)
Vouchers cited: Hume in 1883 (K), Stoddart 875 (K, US), Stoddart 764 (K, US), Stoddart 871 (K, US), Rhyne 683 (US)
British Indian Ocean Territory
Chagos Archipelago
Diego Garcia Island introduced
invasive
Rivers, Julie (2004) (p. 2)
var. velutina
Maldives
Maldive Islands
Addu Atoll   Spicer, R. A./Newberry, D. MacC. (1979) (p. 13)
Maldives
Maldive Islands
Malè Atoll   Fosberg, F. R. (1957) (p. 17)
Mauritius
Mautitius Islands (Mauritius and Rodrigues)
Mauritius Island   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (pp. 11, 17)
French Territory of Mayotte
Mayotte Islands
Mayotte Island introduced
invasive
Comité français de l'Union Internationale pour la Conservation de la Nature en France (2013)
Seychelles
Seychelles Islands
Agalega Island native
Fosberg, F. R./Sachet, Marie-Hélène/Stoddart, D. R. (1983) (p. 110)
Var. mollis (Moq.) Townsend. Vouchers cited: Stoddart 7237 (US)
Seychelles
Seychelles Islands
Alphonse Island   Robertson, I. A. D./Robertson, S. A./Fosberg, F. R. (1983) (p. 179)
Voucher cited: Robertson 2866 (K)
Seychelles
Seychelles Islands
Coetivy Island   Robertson, S. A./Fosberg, F. R. (1983) (p. 146)
Seychelles
Seychelles Islands
Cousin Island native
Fosberg, F. R. (1983) (p. 24)
Voucher cited: Fosberg 52173 (US)
Seychelles
Seychelles Islands
Frégate Island   Robertson, S. A./Todd, D. M. (1983) (p. 44)
Voucher cited: Todd 44
Seychelles
Seychelles Islands
Marie-Louise Island   Wilson, J. R. (1983) (p. 188)
Seychelles
Seychelles Islands
Platte Island   Robertson, S. A./Fosberg, F. R. (1983) (p. 159)
Voucher cited: Robertson 3021 (K)
Seychelles
Seychelles Islands
Poivre Island   Robertson, S. A./Fosberg, F. R. (1983) (p. 167)
Voucher cited: Robertson 2344 (K)
Seychelles
Seychelles Islands
St. Joseph Island   Stoddart, D. R./Coe, M. J./Fosberg, F. R. (1979) (p. 44)
Vouchers cited: Gwynne & Wood 1060 (EA), Stoddart 7148 (US)


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This page was created on 1 JAN 1999 and was last updated on 19 FEB 2013.