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Merr., Fabaceae |
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Present on Pacific Islands? yes
Primarily a threat at high elevations? no
Risk assessment results: High risk, score: 10 (Go to the risk assessment)
Common name(s): [more details]
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Carolinian: serepa, soschghi |
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Chamorro: boiffuring, boiffuring, shoshigi, sosigi, sosugi |
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Chinese: tai wan xiang si |
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English: Formosa acacia, Formosan koa |
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Japanese: sosijyu |
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Palauan: ianángi, yanangi |
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Pohnpeian: pilampwoia |
Habit: tree
Description: "Small tree; adult foliage of falcate phyllodes, juvenile and sucker-shoot foliage of bipinnate leaves; trunk up to 1 m thick in very old trees; phyllodes alternate, coriaceous, parallel-curving-veined, 8-10 cm long, narrowed at both ends; flowers yellow, in small globose heads 6-8 mm in diameter; heads 1 or 2 in axil of phyllode; pods few together, linear or somewhat curved, flat or slightly twisted, brown, 5-10 cm long, 1 cm broad or a little more or less, with about 8 seeds; seeds compressed, brown" (Stone, 1970; p. 295).
Description from Flora of China online.
Habitat/ecology: "Mesic to humid areas in both pastures and forests. Forms monotypic stands in pastures and disturbed sites in forests; eliminates ground cover" (Motooka et al., 2003). Grows well in both dry and mesic habitats. Will often resprout after fire. In Hawaii, it was planted for windbreaks on Maui and has since formed monotypic stands at lower elevations and continues to spread up to 1000 m (Randy Bartlett, communication on Aliens list server).
Propagation: Seeds present in the ground can germinate profusely after fire. Spread through forestry and ornamental plantings. Can be reproduced from cuttings.
Native range: A native of the Northern Philippines, naturalized extensively in Taiwan, Okinawa, and Saipan. In Hawai'i, "widely cultivated and now naturalized in dry to mesic, disturbed areas, 3-915 m" (Wagner et al., 1999; p. 641).
Presence:
| Pacific | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Aguijan (Agiguan, Aguihan) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Aguijan (Agiguan, Aguihan) Island |
Bishop Museum (Honolulu) (1952) (voucher ID: BISH 152469)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Aguijan (Agiguan, Aguihan) Island |
Bishop Museum (Honolulu) (1952) (voucher ID: BISH 152470)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Rota Island |
introduced
invasive cultivated |
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane (2000) (p. 6)
Some naturalization noted. |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
introduced
invasive |
Meyer, Jean-Yves (2000) (p. 101) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
introduced
cultivated |
Space, James C./Falanruw, Marjorie (1999) (p. Field notes.) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
Bishop Museum (Honolulu) (1946) (voucher ID: BISH 152468)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
Bishop Museum (Honolulu) (1946) (voucher ID: BISH 567924)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Saipan Island |
Bishop Museum (Honolulu) (1966) (voucher ID: BISH 659659)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
introduced
cultivated |
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 26)
Voucher cited: Fosberg 31427 (US) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Sarigan Island |
Bishop Museum (Honolulu) (1967) (voucher ID: BISH 401441)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands |
Tinian Island |
introduced
invasive |
Meyer, Jean-Yves (2000) (p. 101) |
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
Bishop Museum (Honolulu) (1935) (voucher ID: BISH 152473)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
Bishop Museum (Honolulu) (1980) (voucher ID: BISH 659660)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Federated States of Micronesia
Chuuk Islands |
Weno (Moen) Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1965) (voucher ID: PTBG 23985)
Taxon name on voucher: Acacia confusa |
|
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
|
Meyer, Jean-Yves (2000) (p. 90)
"Potential invader" |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
|
Glassman, S. F. (1952) (p. 74) |
|
Federated States of Micronesia
Pohnpei Islands |
Pohnpei Island |
introduced
cultivated |
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 88) |
|
Federated States of Micronesia
Yap Islands |
Yap (Waqab) Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Federated States of Micronesia
Yap Islands |
Yap (Waqab) Island |
introduced
|
Meyer, Jean-Yves (2000) (p. 90)
"Potential invader" |
|
Guam
Guam Island |
Guam Island |
introduced
cultivated |
Stone, Benjamin C. (1970) (p. 295) |
|
Guam
Guam Island |
Guam Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Guam
Guam Island |
Guam Island |
Bishop Museum (Honolulu) (1965) (voucher ID: BISH 659661)
Taxon name on voucher: Acacia confusa Merr. |
|
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Guam
Guam Island |
Guam Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1988) (voucher ID: PTBG 22715)
Taxon name on voucher: Acacia confusa |
|
|
Guam
Guam Island |
Guam Island |
National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1965) (voucher ID: PTBG 23144)
Taxon name on voucher: Acacia confusa |
|
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State of Hawaii
Hawaiian Islands |
Hawaii (Big) Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
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State of Hawaii
Hawaiian Islands |
Kahoolawe Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
State of Hawaii
Hawaiian Islands |
Kauai Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
State of Hawaii
Hawaiian Islands |
Lānai Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
State of Hawaii
Hawaiian Islands |
Maui Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
State of Hawaii
Hawaiian Islands |
Molokai Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
State of Hawaii
Hawaiian Islands |
Oahu Island |
introduced
invasive cultivated |
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
Japan (offshore islands)
Bonin (Ogasawara) Islands |
Bonin (Ogasawara) Islands |
introduced
invasive |
Toyoda, Takeshi (2003) (p. 281)
Naturalized |
|
Japan (offshore islands)
Bonin (Ogasawara) Islands |
Bonin (Ogasawara) Islands |
introduced
|
Kato, Hidetoshi (2007) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Palau
Palau (main island group) |
Babeldaob Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 84) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
|
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 90) |
|
Palau
Palau (main island group) |
Koror Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 79) |
|
Palau
Palau (main island group) |
Koror Island |
Bishop Museum (Honolulu) (1950) (voucher ID: BISH 152465)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Palau
Palau (main island group) |
Koror Island |
Beleu National Museum (Republic of Palau. Koror.) (1966) (voucher ID: BNM 414)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Palau
Palau (main island group) |
Koror Island |
Beleu National Museum (Republic of Palau. Koror.) (1967) (voucher ID: BNM 413)
Taxon name on voucher: Acacia confusa Merr. |
|
|
Palau
Palau (main island group) |
Malakal Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 79) |
|
Palau
Palau (main island group) |
Ngerkebesang Island |
introduced
cultivated |
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 79) |
|
Philippines
Philippine Islands |
Philippine Islands |
native
|
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
Philippines
Philippine Islands |
Philippine Islands |
native
|
ILDIS Co-ordinating Centre (2011) |
|
Philippines
Philippine Islands |
Philippine Islands |
invasive
|
Merrill, Elmer D. (1923) (pp. 248-249)
On slopes and in open dry forests at low altitudes, very local. |
| Pacific Rim | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
China
China |
China (People's Republic of) |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
China
China |
China (People's Republic of) |
introduced
cultivated |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
"Widely cultivated". |
|
China
China |
Hong Kong |
introduced
cultivated |
Wu, Te-lin (2001) (p. 129) |
|
Indonesia
Indonesia |
Indonesia (Republic of) |
introduced
|
ILDIS Co-ordinating Centre (2011) |
|
Japan
Japan |
Japan (country) |
introduced
|
Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 184) |
|
Japan
Ryukyu Islands |
Ryukyu Islands |
introduced
|
ILDIS Co-ordinating Centre (2011) |
|
Malaysia
Malaysia |
Malaysia (country of) |
introduced
|
ILDIS Co-ordinating Centre (2011) |
|
Singapore
Singapore |
Singapore (Republic of) |
introduced
cultivated |
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. (2009) (p. 9)
Cultivated only |
|
Taiwan
Taiwan |
Taiwan Island |
native
|
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 641) |
|
Taiwan
Taiwan |
Taiwan Island |
native
|
U.S. Dept. Agr., Agr. Res. Serv. (2011) |
|
Taiwan
Taiwan |
Taiwan Island |
introduced
cultivated |
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong (2011)
"Widely cultivated". |
| Indian Ocean | |||
|
Country/Terr./St. & Island group |
Location |
Cited status &
Cited as invasive & Cited as cultivated & Cited as aboriginal introduction? |
Reference &
Comments |
|
Mauritius
Mautitius Islands (Mauritius and Rodrigues) |
Mauritius Island |
introduced
|
ILDIS Co-ordinating Centre (2011) |
|
Seychelles
Seychelles Islands |
Frégate Island |
Robertson, S. A./Todd, D. M. (1983) (p. 51)
Voucher cited: Procter 4172 |
|
|
Seychelles
Seychelles Islands |
Seychelles Islands |
introduced
|
ILDIS Co-ordinating Centre (2011) |
Comments: This species is being planted in Micronesia. It is a major pest elsewhere. While little reproduction was noted in planted stands, the species should be monitored for invasive behavior. It would probably be wise not to introduce the species to islands where it is not already present (Space & Falanruw, 1999).
An older (probably pre-war) tree was observed in limestone forest on Saipan with no reproduction and a native forest understory, which would indicate limited invasiveness (Space & Falanruw, 1999). Some naturalizing was noted on Rota (Space et al., 2000).
The species is planted in Palau, but no naturalization was noted (Space et al., 2003).
Control:
Chemical: "Saplings sensitive to foliar application of 2,4-D at 1 lb/acre and of triclopyr at 1 lb/acre in a trial at Wailua, Kauai. Triclopyr at 2 lb/acre provided 80% control but 2,4-D at 2 lb/acre caused moderate injury and metsulfuron at 0.4 oz active/acre was ineffective at Kala'e, Molokai. Somewhat tolerant of dicamba. Sensitive to cut-surface applications of 2,4-D, dicamba, glyphosate, picloram (45) and triclopyr in trials at Wailua, Kauai. However, in trials at Kala'e, Molokai, results were less impressive with picloram and dicamba providing serious but not lethal injury and glyphosate and 2,4-D providing moderate injury to Formosan koa. Sensitive to very-low volume basal bark application of 20% triclopyr in oil. Staff at the Hawaii Volcanoes National Park (HAVO) used triclopyr ester at 5% of product in diesel oil applied to the basal bark (Chris Zimmer, HAVO). Tolerant of tebuthiuron pellets." (Motooka et al., 2003)
Additional information:
Excerpt from the book "Weeds
of Hawaiis Pastures and Natural Areas; An Identification and Management Guide"
(Motooka et al., 2003). (PDF format).
Fact sheet from "Common forest trees of Hawaii" (PDF format).
Information from the Global
Invasive Species Database.
Additional online information about Acacia confusa is available from the Hawaiian Ecosystems at Risk project (HEAR).
Information about Acacia confusa as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).
Taxonomic information about Acacia confusa may be available from the Germplasm Resources Information Network (GRIN).
References:
Beleu National Museum (Republic of Palau. Koror.). 1966. Voucher specimen #BNM414(Bechesrrak, M. E-127).
Beleu National Museum (Republic of Palau. Koror.). 1967. Voucher specimen #BNM413(Emmons, B. 3).
Bishop Museum (Honolulu). 1935. Voucher specimen #BISH 152473 (Takamatsu, M. 213).
Bishop Museum (Honolulu). 1946. Voucher specimen #BISH 152468 (Hosaka, E.Y. 2958).
Bishop Museum (Honolulu). 1946. Voucher specimen #BISH 567924 (Hosaka, E.Y. 2958).
Bishop Museum (Honolulu). 1950. Voucher specimen #BISH 152465 (Fosberg, F.R. 32513).
Bishop Museum (Honolulu). 1952. Voucher specimen #BISH 152469 (Kondo, Y.; collector's voucher number unknown).
Bishop Museum (Honolulu). 1952. Voucher specimen #BISH 152470 (Kondo, Y.; collector's voucher number unknown).
Bishop Museum (Honolulu). 1965. Voucher specimen #BISH 659661 (Evans, M. 685).
Bishop Museum (Honolulu). 1966. Voucher specimen #BISH 659659 (Evans, M. 2298).
Bishop Museum (Honolulu). 1967. Voucher specimen #BISH 401441 (Evans, M. 2443).
Bishop Museum (Honolulu). 1980. Voucher specimen #BISH 659660 (Fosberg, F.R. 60366).
Chong, Kwek Yan/Tan, Hugh T. W./Corlett, Richard T. 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species. Raffles Museum of Biodiversity Research, National University of Singapore. 273 pp.
Falanruw, Marjorie C./Maka, Jean E./Cole, Thomas G./Whitesell, Craig D. 1990. Common and scientific names of trees and shrubs of Mariana, Caroline, and Marshall Islands. Pac. SW Forest and Range Expt. Stn. Resource Bulletin PSW-67. 91 pp.
Falanruw, Marjorie C./Maka, Jean E./Cole, Thomas G./Whitesell, Craig D. 1990. Unpublished database and supplement to: Common and scientific names of trees and shrubs of Mariana, Caroline, and Marshall Islands. Pac. SW Forest and Range Expt. Stn. Resource Bulletin PSW-67. 91 pp.
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. 1975. Vascular flora of the northern Marianas Islands. Smithsonian Contrib. Bot. 22:1-45.
Fosberg, F. R./Otobed, D./Sachet, M.-H./Oliver, R. L./Powell, D. A./Canfield, J. E. 1980. Vascular plants of Palau with vernacular names. Department of Botany, The Smithsonian Institution, Washington, D.C. 43 pp.
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce. 1979. A geographical checklist of the Micronesian dicotyledonae. Micronesica 15:1-295.
Glassman, S. F. 1952. The flora of Ponape. Bishop Mus. Bull. 209: 1-152.
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. 2010. Checklist of the vascular plants of Pohnpei with local names and uses. Allertonia, in press. National Tropical Botanical Garden, Lawai, Hawaii. 146 pp.
ILDIS Co-ordinating Centre. 2011. International Legume Database & Information Service. Online searchable database.
Kato, Hidetoshi. 2007. Herbarium records of Makino Herbarium, Tokyo Metropolitan University. Personal communication.
Little, Elbert L./Skolmen, Roger G. 1989. Common forest trees of Hawaii (native and introduced). USDA Agriculture Handbook 679. Washington, D.C. 377 pp. + plates.
Merrill, Elmer D. 1923. An enumeration of Philippine flowering plants, vol. 2 [reprint]. Bureau of Printing, Manila. 530 pp.
Meyer, Jean-Yves. 2000. Preliminary review of the invasive plants in the Pacific islands (SPREP Member Countries). In: Sherley, G. (tech. ed.). Invasive species in the Pacific: A technical review and draft regional strategy. South Pacific Regional Environment Programme, Samoa. 190 pp.
Mito, Toshikazu/Uesugi, Tetsuro. 2004. Invasive alien species in Japan: the status quo and the new regulation for prevention of their adverse effects. Global Environmental Research 8(2)/2004: 171-191.
Motooka, Philip/Castro, Luisa/Nelson, Duane/Nagai, Guy/Ching, Lincoln. 2003. Weeds of Hawaiis Pastures and Natural Areas; An Identification and Management Guide. College of Tropical Agriculture and Human Resources, University of Hawaii at Manoa. 184 pp.
Motooka, Philip/Ching, Lincoln/Nagai, Guy. 2002. Herbicidal Weed Control Methods for Pasture and Natural Areas of Hawaii. Cooperative Extension Service, College of Tropical Agriculture and Human Resources, University of Hawaii. CTAHR free
publication WC-8.
Raulerson, L./Rinehart, A. 1991. Trees and shrubs of the northern Mariana Islands. Saipan: Coastal Resources Management Office of the Governor, Commonwealth of the Northern Mariana Islands 96950. 119 pp.
Robertson, S. A./Todd, D. M. 1983. Vegetation of Frégate Island, Seychelles. In: Sachet, M. H., D. R. Stoddart, and F. R. Fosberg. Floristics and ecology of Western Indian Ocean islands. Atoll Research Bulletin No. 273. Smithsonian Institution, Washington. 253 pp.
Shine, C./Reaser, J. K./Gutierrez, A. T., eds. 2003. Invasive alien species in the Austral-Pacific Region: National Reports & Directory of Resources. Global Invasive Species Programme, Cape Town, South Africa. 185 pp.
Space, James C./Falanruw, Marjorie. 1999. Observations on invasive plant species in Micronesia. USDA Forest Service, Honolulu. Report to the Pacific Islands Committee, Council of Western State Foresters. USDA Forest Service, Honolulu. 32 pp.
Space, James C./Waterhouse, Barbara/Denslow, Julie S./Nelson, Duane. 2000. Invasive plant species on Rota, Commonwealth of the Northern Mariana Islands. USDA Forest Service, Honolulu. 31 pp.
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar. 2003. Report to the Republic of Palau on invasive plant species of environmental concern. USDA Forest Service, Honolulu. 174 pp.
Stone, Benjamin C. 1970. The flora of Guam. Micronesica 6:1-659.
Toyoda, Takeshi. 2003. Flora of the Bonin Islands. Aboc & Co. 522 pp.
U. S. Government. 2011. Integrated Taxonomic Information System (ITIS) (on-line resource).
U.S. Dept. Agr., Agr. Res. Serv. 2011. National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Online searchable database.
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. 1999. Manual of the flowering plants of Hawaii. Revised edition. Bernice P. Bishop Museum special publication. University of Hawaii Press/Bishop Museum Press, Honolulu. 1919 pp. (two volumes).
Wu, Te-lin. 2001. Check List of Hong Kong Plants. Hong Kong Herbarium and the South China Institute of Botany. Agriculture, Fisheries and Conservation Department Bulletin 1 (revised). 384 pp.
Zhengyi, Wu/Raven, Peter H./Deyuan, Hong. 2011. Flora of China (online resource).