Herbst, Derral R./Wagner, Warren L. 1999. Contributions to the flora of Hawaii. VII. In: Evenhuis, Neal L. and Eldredge, Lucius G., eds. Records of the Hawaii Biological Survey for 1998. Part 1: Articles. Bishop Museum Occasional Papers. 58:12-36.
Online details: http://hbs.bishopmuseum.org/pdf/herbst&wagner99.pdf