Pacific Island Ecosystems at Risk (PIER)

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Physalis angulata
L., Solanaceae
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Present on Pacific Islands?  yes

Primarily a threat at high elevations?  no

Other Latin names:  Physalis lanceifolia Nees

Common name(s): [more details]

Chamorro: tomate chaca, tomates caputi

Chinese: ku zhi

English: annual ground cherry, bladder cherry, bladderberry, cape gooseberry, cutleaf groundcherry, ground cherry, wild cape gooseberry

Fijian: cevucevu, kospeli, mborosousou, mborosousou ni vavalangi, mbotembote yandra, mburasu, mokoai, thevuthevu

I-Kiribati: te baraki, te bin

Japanese: sennari-houzuki, sennari-hozuki

Maori (Cook Islands): tāmaru, tāmaru, tūpera tāmaru-ariki, tūpere

Marshallese: kaõrõr, kaoror

Nauruan: oatamo, watamo

Niuean: manini

Palauan: bubeubedul, pohaa

Pohnpeian: tuhke wah pwonopwon duwehte tehn sele

Samoan: māgalo, māgalo, vī vao, vī vao

Spanish: bolsa mullaca, bomba, capuli cimarron, huevillo, huevo de tortuga, meltonate, mullaca, popoja, tomatillo, vejigon

Tahitian: tamanufairi, tamaru ha‘ari

Tongan: polo pā

Tuvaluan: pini, te peen

Habit:  herb

Description:  "Branched erect nearly glabrous herb; stems angled, hollow, leaves ovate to ovate-oblong, obtuse or acute or rounded (somewhat asymmetric) at base, acute-acuminate, often repand-dentate, rather dark dull green, very short-puberulent on both sides on the nerves, 3-15 cm long, 2.5-10 cm wide, on long petioles 2-10 cm, calyx 4-6 mm long in flower, 2-4 long in fruit, short-puberulent; corolla nearly 1 cm long, yellowish-green, partly puberulent externally; anthers blue, 1-2 mm; fruit about 1-1.8 cm long, ellipsoid, within the balloon-like calyx (which has a small apical opening); seeds about 1.7 mm long, flat, yellowish, reniform"  (Stone, 1970; p. 520).

Habitat/ecology:  "It infests primarily annual crops but also occurs in orchards, nurseries, fallow land, roadsides and waste areas. It grows best in moist, fertile soils and is tolerant of partial shade"  (Holm et al., 1997; pp. 580, 582).  "Common to abundant in croplands and disturbed places" (Whistler, 1988; p. 36).

In Hawai‘i, "sparingly naturalized in disturbed sites, ca. 90 m" (Wagner et al., 1999; p. 1265).  In Fiji, "often locally abundant as a weed in villages, waste places, pastures, plantations, and canefields, along roads, on open slopes, and in forest along creeks from near sea level to about 600 m" (Smith, 1991; pp. 29-30).  In Niue, "a fairly common to very common weed of plantations in many areas, particularly the Basin" (Sykes, 1970; p. 196).  In New Guinea, "a weed of gardens, waste land and cultivated fields. From low altitudes to about 1000 m" (Henty & Pritchard, 1975; p. 149). In Tonga, "an occasional plantation weed" (Yuncker, 1959; p. 236).

Propagation:  Seed

Native range:  Possibly from tropical America, now a common tropical weed (Stone, 1970; p. 520).  North America to Central America (Wagner et al., 1999; p. 1265).  "An American species apparently aboriginally introduced to Polynesia" (Whistler, 1988; p. 36).

Presence:

Pacific
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
American Samoa
Manu‘a Islands
Ta‘u Island   Swarbrick, John T. (1997) (p. 95)
American Samoa
Manu‘a Islands
Ta‘u Island   Whistler, W. A. (1990)
Vouchers cited: Whistler 1300, Whistler 3330
American Samoa
Manu‘a Islands
Ta‘u Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1990) (voucher ID: PTBG 11079)
Taxon name on voucher: Physalis angulata
American Samoa
Swain's Island
Swain's Island introduced
invasive
Whistler, W. A. (1983) (p. 15)
Voucher cited: Whistler 3439
American Samoa
Tutuila Islands
Tutuila Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1992) (voucher ID: PTBG 44177)
Taxon name on voucher: Physalis angulata
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Anatahan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Anatahan Island introduced
invasive
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1975) (p. 39)
Voucher cited: Falanruw 1650 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Farallon de Medinilla (Marpi, Medinilla) Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Pagan Island introduced
Fosberg, F. R./Falanruw, M. V. C./Sachet, M.-H. (1980) (p. 213)
Voucher cited: Falanruw 3318 (US)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Rota Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Also var. lanceifolia (Nees) Waterfall
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Rota Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1978) (voucher ID: PTBG 32926)
Taxon name on voucher: Physalis angulata
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Saipan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
var. lanceifolia (Nees) Waterfall
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Sarigan Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Commonwealth of the Northern Mariana Islands (US)
Northern Mariana Islands
Tinian Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Also var. lanceifolia (Nees) Waterfall
Cook Islands
Northern Cook Islands
Suwarrow (Taka) Atoll   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
‘Atiu Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Mangaia Island   McCormack, Gerald (2013)
Cook Islands
Southern Cook Islands
Ma‘uke Island   McCormack, Gerald (2013)
Ecuador (Galápagos Islands)
Española Group
Española Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Española Group
Gardner por Espaņola Islet, Española Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Española Group
Oeste Islet, Española Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Española Group
Osborn Islet, Española Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Floreana Group
Campéon Islet, Floreana Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Floreana Group
Floreana Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Isabela Group
Volcán Sierra Negra, Isabela Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Pinta Group
Pinta Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Edeén Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Mosquera Islet, Santa Cruz Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Plaza Norte Islet, Santa Cruz Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Punta Bowditch Norte Islet, Santa Cruz Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Punta Bowditch Sur Islet, Santa Cruz Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Santa Cruz Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Cruz Group
Seymour Norte Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santa Fé Group
Santa Fé Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santiago Group
Santiago Island native
Charles Darwin Research Station (2005)
Ecuador (Galápagos Islands)
Santiago Group
Sombrero Chino Islet, Santiago Island native
Charles Darwin Research Station (2005)
Federated States of Micronesia
Chuuk Islands
Chuuk Islands introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Federated States of Micronesia
Chuuk Islands
Pis Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Federated States of Micronesia
Chuuk Islands
Weno (Moen) Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Federated States of Micronesia
Kosrae Island
Kosrae Island introduced
invasive
Lorence, David H./Flynn, Timothy (2010) (p. 24)
Voucher cited: Lorence 9394 (PTBG)
Naturalized
Federated States of Micronesia
Kosrae Island
Kosrae Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1996) (voucher ID: PTBG 24905)
Taxon name on voucher: Physalis angulata L.
Federated States of Micronesia
Kosrae Island
Kosrae Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2005) (voucher ID: PTBG 44961)
Taxon name on voucher: Physalis angulata
Federated States of Micronesia
Pohnpei Islands
Ant Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Also var. lanceifolia (Nees) Waterfall
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island introduced
invasive
Herrera, Katherine/Lorence, David H./Flynn, Timothy/Balick, Michael J. (2010) (p. 137)
Voucher cited: R. Gallen 149 (NY, PTBG)
Naturalized
Federated States of Micronesia
Pohnpei Islands
Pohnpei Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 43522)
Taxon name on voucher: Physalis angulata
Federated States of Micronesia
Yap Islands
Fais Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Federated States of Micronesia
Yap Islands
Fais Island introduced
invasive
Fosberg, F. R./Evans, M. (1969) (p. 13)
Voucher cited: Fosberg 46679 (US)
Federated States of Micronesia
Yap Islands
Faraulep (Faraulap) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Federated States of Micronesia
Yap Islands
Ulithi Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Federated States of Micronesia
Yap Islands
Yap (Waqab) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Fiji
Fiji Islands
Taveuni Island introduced
invasive
Smith, Albert C. (1991) (pp. 29-31)
Voucher cited: DA 8926
Fiji
Fiji Islands
Vanua Levu Island introduced
invasive
Smith, Albert C. (1991) (pp. 29-31)
Voucher cited: DA 10473
Fiji
Fiji Islands
Viti Levu Island introduced
invasive
Smith, Albert C. (1991) (pp. 29-31)
Vouchers cited: Greenwood 188, Degener & Ordonez 13688, DA 9695, Smith 4442, DA 9487, Smith 9445, DA 9553, DA 10046, DA 9568, DA 11086, DA 9971, DA 10603, DA 3169
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Fatu Hiva Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Fatu Hiva Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1974) (voucher ID: PTBG 8907)
Taxon name on voucher: Physalis angulata
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Hiva Oa Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Hiva Oa Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (2003) (voucher ID: PTBG 39749)
Taxon name on voucher: Physalis angulata
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Nuku Hiva (Nukahiva) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 4108 (PAP), J. Florence 4172 (PAP), J. Florence 8442 (PAP)
Adventice
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Tahuata (Tuhuata) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Ua Huka (Huahuna, Uahuka) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Lorence, David H./Wagner, Warren L. (2013)
French Polynesia
Marquesas Islands
Ua Pou (Huapu, Uapou, Uapu) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 4446 (PAP)
Adventice
French Polynesia
Society Islands
Mehetia (Mahetia, Meetia) Island   Welsh, S. L. (1998)
French Polynesia
Society Islands
Mehetia (Mahetia, Meetia) Island   Fosberg, F. R. (1997) (p. 143)
French Polynesia
Society Islands
Moorea Island   Welsh, S. L. (1998)
French Polynesia
Society Islands
Moorea Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 2171 (PAP)
Adventice
French Polynesia
Society Islands
Raiatea (Havai) Island   Welsh, S. L. (1998)
French Polynesia
Society Islands
Raiatea (Havai) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 3384 (PAP)
Adventice
French Polynesia
Society Islands
Taha‘a Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 10591 (PAP)
Adventice
French Polynesia
Society Islands
Tahiti Island   Welsh, S. L. (1998)
French Polynesia
Society Islands
Tahiti Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 3201 (PAP), F.R. Fosberg 65013 (PAP)
Adventice
French Polynesia
Society Islands
Tupai Island introduced
invasive
Sachet, M.-H. (1983) (p. 22)
Voucher cited: Sachet 1975 (US)
French Polynesia
Society Islands
Tupai Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 1975 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Hao (Hau) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Tuamotu Archipelago
Niau Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 10111 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Rangiroa (Rairoa) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 10004 (PAP)
Adventice
French Polynesia
Tuamotu Archipelago
Takapoto Atoll introduced
invasive
Sachet, M-H. (1983) (p. 36)
Voucher cited: Sachet 2052 (US)
French Polynesia
Tuamotu Archipelago
Tikehau (Tikahau) Atoll introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 7079 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Raivavae (Raevavae) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Voucher cited: J. Florence 5819 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rimatara (Rimitara) Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Adventice
French Polynesia
Austral (Tubuai) Islands
Rurutu Island introduced
invasive
Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. (2013)
Vouchers cited: J. Florence 6140 (PAP), J. Florence 6164 (PAP)
Adventice
French Polynesia
Austral (Tubuai) Islands
Tubuai Island   Welsh, S. L. (1998)
Guam
Guam Island
Guam Island introduced
invasive
Stone, Benjamin C. (1970) (p. 520)
Voucher cited: Stone 4314 (GUAM). Also var. lanceifolia, listed as Physalis lanceifolia Nees.; vouchers cited: McGregor 512, G.E.S. 202, Fosberg 43448, Anderson 22, Hosaka 3190.
Guam
Guam Island
Guam Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Also var. lanceifolia (Nees) Waterfall
State of Hawaii
Hawaiian Islands
Hawai‘i (Big) Island introduced
invasive
Oppenheimer, Hank L. (2003) (p. 26)
Voucher cited: Oppenheimer H80106 (BISH, PTBG)
State of Hawaii
Hawaiian Islands
Kaua‘i Island introduced
invasive
Wagner, Warren L./Herbst, Derral R./Sohmer, S. H. (1999) (p. 1265)
Voucher cited: Herbst & Ishikawa 5664 (BISH)
State of Hawaii
Hawaiian Islands
Moloka‘i Island introduced
invasive
Staples, George W./Herbst, Derral R./Imada, Clyde T. (2006) (p. 8)
Voucher cited: V. Caraway 150 (BISH)
State of Hawaii
Hawaiian Islands
O‘ahu Island introduced
invasive
Imada, Clyde T./Staples, George W./Herbst, Derral R. (2000) (p. 15)
Voucher cited: R. Heu & W. Nagamine s.n. (BISH 658032)
Kiribati
Line Islands
Kiritimati (Christmas) Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1995) (voucher ID: PTBG 38220)
Taxon name on voucher: Physalis angulata L.
Kiribati
Phoenix Islands
Enderbury Island introduced
invasive
Fosberg, F. R./Stoddart, D. R. (1994) (pp. 48-49)
Note seen in 1975. Voucher cited: Clegern 168 (BISH)
Kiribati
Phoenix Islands
Kanton Island introduced
invasive
Fosberg, F. R./Stoddart, D. R. (1994) (pp. 48-49)
Vouchers cited: Fosberg & Stoddart 54894 (US, HAW, K), Degener & Hatheway 21411 (BISH), Fosberg 55803 (US, BISH), Degener & Degener 24654 (BISH)
Kiribati
Phoenix Islands
Orona (Hull) Atoll introduced
invasive
Fosberg, F. R./Stoddart, D. R. (1994) (pp. 48-49)
Vouchers cited: Fosberg & Stoddart 54701 (US, HAW, K) (but may be P. lanceifolia Nees).
Kiribati
Tungaru (Gilbert) Islands
Abaiang Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
also var. lanceifolia (Nees) Waterfall
Kiribati
Tungaru (Gilbert) Islands
Abemama Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 76)
Voucher: Imada 2003-166 (BISH 697777)
Kiribati
Tungaru (Gilbert) Islands
Butaritari Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
also var. lanceifolia (Nees) Waterfall
Kiribati
Tungaru (Gilbert) Islands
Butaritari Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 77)
Voucher: Imada 2003-76 (BISH 697691)
Kiribati
Tungaru (Gilbert) Islands
Butaritari Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1972) (voucher ID: PTBG 32928)
Taxon name on voucher: Physalis angulata
Kiribati
Tungaru (Gilbert) Islands
Maiana Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 79)
Voucher: Imada 2003-113 (BISH 697724)
Kiribati
Tungaru (Gilbert) Islands
Nonouti Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
also var. lanceifolia (Nees) Waterfall
Kiribati
Tungaru (Gilbert) Islands
Onotoa Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
also var. lanceifolia (Nees) Waterfall
Kiribati
Tungaru (Gilbert) Islands
Onotoa Atoll   Moul, Edwin T. (1957) (p. 21)
Vouchers cited: Moul 8093 (US), Moul 8149 (US)
Kiribati
Tungaru (Gilbert) Islands
Tarawa Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
also var. lanceifolia (Nees) Waterfall
Kiribati
Tungaru (Gilbert) Islands
Tarawa Atoll introduced
invasive
Space, James C./Imada, Clyde T. (2004) (p. 75)
Voucher: Imada 2003-21 (BISH 697636)
Kiribati
Tungaru (Gilbert) Islands
Tarawa Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1972) (voucher ID: PTBG 32927)
Taxon name on voucher: Physalis angulata
Marshall Islands
Ralik Chain
Ailinglaplap (Ailinglapalap) Atoll   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Marshall Islands
Ralik Chain
Bikini (Pikinni) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Marshall Islands
Ralik Chain
Bikini (Pikinni) Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1985) (voucher ID: PTBG 23939)
Taxon name on voucher: Physalis angulata
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll introduced
invasive
Lamberson, J. O. (1982) (p. 56)
Marshall Islands
Ralik Chain
Enewetak (Ãnewetak, Eniwetok) Atoll introduced
invasive
St. John, Harold (1960) (p. 333)
Vouchers cited: Lane 57-139, Lane 57-101
Marshall Islands
Ralik Chain
Jaluit (Jãlwõj) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Marshall Islands
Ralik Chain
Jaluit (Jãlwõj) Atoll introduced
invasive
Fosberg, F. R./Sachet, M.-H. (1962) (p. 34)
Voucher cited: Fosberg 26710 (US)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
invasive
Whistler, W. A./Steele, O. (1999) (pp. 67-68)
Vouchers cited: Fosberg 48053, Fosberg 48054, Herbst 8948, Herbst 9619, Herbst 9623, Herbst 9624, Whistler 11168, Whistler 11194, Whistler 11261
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 19)
Voucher cited: Fosberg 34381 (US)
Marshall Islands
Ralik Chain
Kwajalein (Kuwajleen) Atoll   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1998) (voucher ID: PTBG 44984)
Taxon name on voucher: Physalis angulata
Marshall Islands
Ralik Chain
Ujae Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Marshall Islands
Ralik Chain
Ujae Atoll introduced
invasive
Fosberg, F. R. (1955) (p. 19)
Voucher cited: Fosberg 34286 (US)
Marshall Islands
Ratak Chain
Arno Atoll introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Marshall Islands
Ratak Chain
Arno Atoll   Anderson, Donald (1951) (p. v)
Voucher cited: Anderson 3652
Marshall Islands
Ratak Chain
Arno Atoll introduced
Vander Velde, Nancy/Vander Velde, Brian (2007) (p. 12)
"Uncommon; few plants in ruderal areas of Arno and Matlon villages".
Marshall Islands
Ratak Chain
Jemo Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Marshall Islands
Ratak Chain
Jemo Island introduced
invasive
Fosberg, F. R. (1955) (p. 19)
Voucher cited: Fosberg 33872 (US)
Marshall Islands
Ratak Chain
Majuro (Mãjro) Atoll introduced
invasive
Vander Velde, Nancy (2003) (p. 127)
Marshall Islands
Ratak Chain
Maloelap Atoll   Vander Velde, Nancy (year unknown)
Marshall Islands
Ratak Chain
Mejit (Mãjej) Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Marshall Islands
Ratak Chain
Mili (Mile) Atoll introduced
invasive
Vander Velde, Nancy/Vander Velde, Brian (2010) (p. A22)
Uncommon; in ruderal areas, Mili, Arbar, Tokewa, Bar. Voucher 0723201001 (BISH, GUAM, PTBG)
Marshall Islands
Ratak Chain
Wotje (Wõjjã) Atoll introduced
Vander Velde, Nancy/Vander Velde, Brian (2009) (p. 2)
Voucher cited: Vander Velde 0514200929
Nauru
Nauru Island
Nauru Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Nauru
Nauru Island
Nauru Island introduced
invasive
Thaman, R. R./Fosberg, F. R./Manner, H. I./Hassall, D. C. (1994) (pp. 198-199)
Vouchers cited: Fosberg 58645 (US), Fosberg 58768 (US), Scully 130N (US), Thaman & Manner 14 (SUVA), Thaman & Manner 22321 (SUVA), Swarbrick 9541 (BRIU)
New Caledonia
New Caledonia
New Caledonia Islands   Swarbrick, John T. (1997) (p. 95)
New Caledonia
New Caledonia
New Caledonia Islands introduced
invasive
Tassin, Jacques (2005)
New Caledonia
New Caledonia Archipelago
Île Grande Terre introduced
invasive
MacKee, H. S. (1994) (p. 131)
Vouchers cited: MacKee 28350, MacKee 29508
Spontané
Niue
Niue
Niue Island introduced
invasive
Sykes, W. R. (1970) (p. 196)
Voucher cited: CHR 169998
Niue
Niue
Niue Island introduced
invasive
abo. intro.
Whistler, W. A. (1988) (p. 36)
Niue
Niue
Niue Island introduced
invasive
abo. intro.
Whistler, W. A./Atherton, J. (1997) (p. 46)
Probably a Polynesian introduction.
Niue
Niue
Niue Island introduced
invasive
abo. intro.
Space, James C./Waterhouse, Barbara M./Newfield, Melanie/Bull, Cate (2004) (p. 50)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 104)
Palau
Palau (Belau ) (main island group)
Angaur (Ngeaur) Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Palau
Palau (Belau ) (main island group)
Babeldaob Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 102)
Palau
Palau (Belau ) (main island group)
Babeldaob Island introduced
invasive
Space, James C./Lorence, David H./LaRosa, Anne Marie (2009) (p. 64)
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Palau
Palau (Belau ) (main island group)
Koror (Oreor) Island   Beleu National Museum (Republic of Palau. Koror.) (1966) (voucher ID: BNM 752)
Taxon name on voucher: Physalis angulata L.
Palau
Palau (Belau ) (main island group)
Malakal (Ngemelachel) Island introduced
Fosberg, F. R./Sachet, Marie-Hélène/Oliver, Royce (1979) (p. 245)
Palau
Palau (Belau ) (main island group)
Malakal (Ngemelachel) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau
Palau (Belau ) (main island group)
Malakal (Ngemelachel) Island   Beleu National Museum (Republic of Palau. Koror.) (1968) (voucher ID: BNM 1406)
Taxon name on voucher: Physalis angulata L.
Palau
Palau (Belau ) (main island group)
Ngerkebesang Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 98)
Palau
Palau (Belau ) (main island group)
Peleliu (Beliliou) Island introduced
invasive
Space, James C./Waterhouse, Barbara/Miles, Joel E./Tiobech, Joseph/Rengulbai, Kashgar (2003) (p. 106)
Palau
Sonsorol Group
Pulo Ana Island   Wagner, W. L./ Herbst, D. R./Weitzman, A./Lorence, D.H. (2013)
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island) invasive
Henty, E. E./Pritchard, G. H. (1975) (p. 149)
From low altitudes to about 1000 m.
Papua New Guinea
Papua New Guinea (eastern New Guinea Island)
Papua New Guinea (eastern New Guinea Island)   Waterhouse, D. F. (1997) (p. 64)
Philippines
Philippine Islands
Philippine Islands   Waterhouse, D. F. (1993) (p. 68)
Philippines
Philippine Islands
Philippine Islands introduced
invasive
Merrill, Elmer D. (1923) (p. 423)
Widely scattered in open waste places in and near settlements.
Samoa
Western Samoa Islands
Western Samoa Islands   Whistler, W. A. (1988) (p. 36)
Solomon Islands
Solomon Islands
Solomon Islands   Swarbrick, John T. (1997) (p. 95)
Solomon Islands
Solomon Islands
Solomon Islands native
Hancock, I. R./Henderson, C. P. (1988) (p. 99)
Tonga
Ha‘apai Group
Ha‘apai Islands   Whistler, W. A. (1992) (p. 73)
Voucher cited: Soakai 311
Tonga
Ha‘apai Group
Tofua Island   Whistler, W. A. (1992) (p. 73)
Voucher cited: Scarth-Johnson 92
Tonga
Tongatapu Group
‘Eua Island   Whistler, W. A. (1992) (p. 73)
Voucher cited: Soakai 343
Tonga
Tongatapu Group
Tongatapu Island   Whistler, W. A. (1992) (p. 73)
Vouchers cited: Lawrence 104, Soakai 506
Tonga
Tongatapu Group
Tongatapu Island introduced
invasive
Yuncker, T. G. (1959) (p. 236)
Voucher cited: Yuncker 15098
Tonga
Tongatapu Group
Tongatapu Island   National Tropical Botanical Garden (U.S.A. Hawaii. Kalaheo.) (1989) (voucher ID: PTBG 6762)
Taxon name on voucher: Physalis angulata
Tonga
Vava‘u Group
Vava‘u Island   Whistler, W. A. (1992) (p. 73)
Vouchers cited: Harvey s.n., Crosby 123
Tuvalu
Tuvalu (Ellis) Islands
Nui Atoll   Woodroffe, C. D. (1985) (p. 15)
Voucher cited: Woodroffe 145 (CHR)
Tuvalu
Tuvalu (Ellis) Islands
Tuvalu Islands   Swarbrick, John T. (1997) (p. 95)
Vanuatu
New Hebrides Islands
Vanuatu (Republic of)   Swarbrick, John T. (1997) (p. 95)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis (‘Uvea) Island introduced
invasive
Meyer, Jean-Yves (2007) (p. 38)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis and Futuna Islands   Whistler, W. A. (1988) (p. 36)
Wallis and Futuna
Wallis and Futuna (Horne) Islands
Wallis and Futuna Islands introduced
Meyer, Jean-Yves (2000) (p. 107)
"Potential invader". Gardens, secondary vegetation.
Pacific Rim
Country/Terr./St. &
Island group
Location Cited status &
Cited as invasive &
Cited as cultivated &
Cited as aboriginal introduction?
Reference &
Comments
Australia
Australia (continental)
New South Wales introduced
invasive
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Australia
Australia (continental)
Queensland introduced
invasive
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Cambodia
Cambodia
Cambodia (Kingdom of)   Waterhouse, D. F. (1993) (p. 68)
China
China
China (People's Republic of)   Holm, Leroy/Pancho, Juan V./Herberger, James P./Plucknett, Donald L. (1979) (p. 278)
China
China
Hong Kong native
Wu, Te-lin (2001) (p. 219)
In forests of valleys, on roadsides near villages.
Colombia
Colombia
Colombia (Republic of)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Ecuador (Mainland)
Ecuador
Ecuador (Republic of) (continental)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
El Salvador
El Salvador
El Salvador (Republic of)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 584)
Guatemala
Guatemala
Guatemala (Republic of)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 584)
Honduras
Honduras
Honduras (Republic of)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 583)
Indonesia
Indonesia
Indonesia (Republic of)   Waterhouse, D. F. (1993) (p. 68)
Japan
Japan
Japan (country) introduced
invasive
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Japan
Japan
Japan (country) introduced
Mito, Toshikazu/Uesugi, Tetsuro (2004) (p. 188)
var. angulata, var. lanceifolia (Nees) Waterf. and var. pendula (Rydb.) Waterf.
South Korea
South Korea
South Korea (Republic of) introduced
invasive
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Malaysia
Malaysia
Malaysia (country of)   Waterhouse, D. F. (1993) (p. 68)
Mexico
Mexico
Mexico (United Mexican States)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Nicaragua
Nicaragua
Nicaragua (Republic of)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 583)
Perú
Perú
Perú (Republic of)   Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Taiwan
Taiwan Island
Taiwan Island introduced
invasive
Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James (1997) (p. 580)
Taiwan
Taiwan Island
Taiwan Island introduced
invasive
Ching-I Peng, ed. (2013)
"Common in lowland areas of disturbance and cropland".
Thailand
Thailand
Thailand (Kingdom of)   Waterhouse, D. F. (1993) (p. 68)
Vietnam
Vietnam
Vietnam (Socialist Republic of)   Waterhouse, D. F. (1993) (p. 68)

Comments:  See also Physalis minima.

Additional information:
Additional online information about Physalis angulata is available from the Hawaiian Ecosystems at Risk project (HEAR).

Information about Physalis angulata as a weed (worldwide references) may be available from the Global Compendium of Weeds (GCW).

Taxonomic information about Physalis angulata may be available from the Germplasm Resources Information Network (GRIN).

References:

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Beleu National Museum (Republic of Palau. Koror.). 1966. Voucher specimen #BNM752(Cheatham 138).

Beleu National Museum (Republic of Palau. Koror.). 1968. Voucher specimen #BNM1406(Blackburn, B.S. 178).

Charles Darwin Research Station. 2005. CDRS Herbarium records.

Ching-I Peng, ed. 2013. Digital flora of Taiwan (online resource).

Florence, J./Chevillotte, H./Ollier, C./Meyer, J.-Y. 2013. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP). (online resource).

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Zhengyi, Wu/Raven, Peter H./Deyuan, Hong. 2013. Flora of China (online resource).


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This page was created on 26 JUL 2002 and was last updated on 22 JAN 2007.